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Arthropoda Selecta 11 (4): 283317

ї ARTHROPODA SELECTA, 2002

The spider genus Clubiona Latreille, 1804 (Aranei: Clubionidae) in the fauna of the former USSR: 2003 update Пауки рода Clubiona Latreille, 1804 (Aranei: Clubionidae) фауны бывшего СССР: обновление 2003 года K.G. Mikhailov К.Г. Михайлов
Zoological Museum, Moscow Lomonosov State University, Bolshaya Nikitskaya Str. 6, Moscow 125009 Russia. Зоологический музей МГУ, Большая Никитская ул. 6, Москва 125009 Россия.

KEY WORDS: spiders, Clubiona, taxonomy, faunistics, former USSR. КЛЮЧЕВЫЕ СЛОВА: пауки, Clubiona, систематика, фаунистика, бывший СССР. ABSTRACT: Additional faunistic data for the spider genus Clubiona Latreille, 1804 in the former USSR are provided. In total, 91 species (one undescribed) are considered. All the literature-derived data since Mikhailov [1992b] are included. The following new species are described: C. mazandaranica sp.n. (, Iran: Mazandaran Prov., Azerbaijan: Lenkoran Prov.), C. mykolai sp.n. (, Ukraine: The Crimea), C. yaroslavi sp.n. (), C. dunini sp.n. (), C. helenae sp.n. (), C. sichotanica sp.n. () (all from Russia: Maritime Prov. = Primorie). C. congentilis Kulczyсski, 1913 () is figured and redescribed. The female of C. hyrcanica Michailov, 1990 and the male of C. pseudoneglecta Wunderlich, 1994 are figured and described for the first time. Deviations in or details of the copulatory organs of C. latericia Kulczyсski, 1926 and C. microsapporensis Michailov, 1990 are figured. The following new synonymies are established: C. liachviana Mkheidze, 1997 = C. alpicola Kulczyсski, 1881; C. chechtsirica Michailov, 1995 = C. paiki Michailov, 1991; C. yadongensis Hu, 2001 = C. xiningensis Hu, 2001 = C. neglecta O.Pickard-Cambridge, 1862; C. serrulata Hu, Wang et Wang, 1991 = C. wolchongsensis Paik, 1990 = C. papillata Schenkel, 1936. РЕЗЮМЕ: Приведены дополнительные данные по фауне пауков рода Clubiona Latreille, 1804 в бывшем СССР. Всего обнаружен 91 вид (в т.ч. один неописанный). Включены все литературные данные, опубликованые после работы Михайлова [Mikhailov, 1992b]. Описаны новые виды: C. mazandaranica sp.n. (, Иран: пров. Мазандаран, Азербайджан: Ленкорань), C. mykolai sp.n. (, Украина: Крым), C. yaroslavi sp.n. (), C. dunini sp.n. (), C. helenae sp.n. (), C. sichotanica sp.n. () (все из России, Приморье). Дано илюстрированное переописание C. congentilis Kulczyсski , 1913 ( ). Впервые описаны самка C. hyrcanica Michailov, 1990 и самец C. pseudoneglecta Wunderlich, 1994. Изображены
Printed in 2003.

отклонения в строении и особенности строения копулятивных органов C. latericia Kulczyсski, 1926 и C. microsapporensis Michailov, 1990. Установлены новые синонимы: C. liachviana Mkheidze, 1997 = C. alpicola Kulczyсski, 1881; C. chechtsirica Michailov, 1995 = C. paiki Michailov, 1991; C. yadongensis Hu, 2001 = C. xiningensis Hu, 2001 =C. neglecta O.PickardCambridge, 1862; C. serrulata Hu, Wang et Wang, 1991 = C. wolchongsensis Paik, 1990 = C. papillata Schenkel, 1936. The aim of this paper is to provide updated faunistic information for the spider genus Clubiona in the exUSSR [Mikhailov, 1992b]. In totaltally, 71 species of Clubiona were known from the ex-USSR in 1992 [Mikhailov, 1992b], 80 in 1995 [Mikhailov, 1995a] and 81 in 1997 [Mikhailov, 1997a]. This paper lists 91 species, including 6 new to science and one yet to be described. A redescription of Clubiona congentilis Kulczyсski, 1913 was omitted in my 1992 paper, because figures were missed at that time; a redescription and figures based on the material cited in Mikhailov [1992b] are provided herein.

To assist proper identification, references to (re)-descriptions and figures are given for Asian, especially Oriental, species. Page numbers in the corresponding papers are not quoted when only geographical data are provided. For nomina dubia and nomina nuda, see the catalogue of the exUSSR spiders [Mikhailov, 1997a]. Unfortunately, a detailed ecological study and precise collecting data within local habitats has not yet been conducted in Maritime Prov. (= Primorie), the Russian Far East. Therefore, many previously described species are known from only one sex. A proper ecological investigation is needed. The following abbreviations are used in the descriptions: Cb cymbium; d. dorsally; EP embolar (embolic) part of bulbus; F femur; lat. laterally; ld literature data; Mt metatarsus; Pt patella; rlat. retrolaterally; T tarsus; TA tibial apophysis; Ti tibia; v. ventrally. Collector names are also abbreviated: AP Alexandr V. Ponomarev; AT Andrei V. Tanasevitch; AZ Andrei S. Zorkin; DL Dmitri V. Logunov; KE Kirill Yu. Eskov;


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TLS, RZN, BRN, ORL, TMB, KUR, BLG, VRN, MRS, ChVSh, ULN, TTR, SMR, AST, RST, KLM, KRS, STV, KBA, ARKh, KOM, KRV, PRM, EKB, ChLB, ORN, KRG, TMN, NVS, TMS, KMR, ALT, KRN, Tuva, YaKT, MGD, KMCh, IRK, BUR, ChIT, AMR, KhAB, PRI, SAKh. ESTONIA. LATVIA. LITHUANIA. (no subdivision) BELARUS. UKRAINE. ChRK Cherkassy Area, DNTs Donetsk Area, KhRK Kharkov Area, KhRS Kherson Area, KRV Kirovograd Area, ODS Odessa Area, SUM Sumy Area, ZAK Zakarpatskaya Area, ZhTM Zhitomir Area. MOLDAVIA (MOLDOVA). AZERBAIJAN. GEORGIA. (no subdivision) KAZAKHSTAN. AKM Akmolinsk Area, PVL Pavlodar Area, SKO Severo-Kazakhstanskaya (=North Kazakhstan) Area, VKO Vostochno-Kazakhstanskaya (=East Kazakhstan) Area. UZBEKISTAN. KIRGHIZIA (KYRGHYZSTAN). (no subdivision) Literature data mainly cover publications since 1992, and those not mentioned in Mikhailov [1992b]. Several earlier contributions from the 1970s that were missed in my paper are also included. Data that consist only of citations by earlier authors are identified using -ld, meaning literature data. Literature data derived from the collections are marked with an asterisk (*). In the Distribution section, data for the territories outside the ex-USSR start as a new paragraph.

KM Kirill G. Mikhailov; NK Nikolai (=Mykola) M. Kovblyuk; OL Oleg V. Lyakhov; PD Peter M. Dunin (); SG Sergei I. Golovatch; SK Seppo Koponen; TO Tatiana I. Oliger; YM Yuri M. Marusik. The typical pattern of leg armature in Clubiona is as follows: F III d.1.1.2, IIIIV d.1.1.3, Pt IIIIV rlat.1, Ti III v.2.2, III d.2.2, v.1.1, IV d.2.2, v.1.1.1, Mt III v.2, III d.2.1.2, lat.1.2, v.2.2, IV d.2.1.2, lat.2.2, v.2.1.2. Only deviations from this formula are given in the descriptions. All measurements are in mm. When several specimens are measured, M+m is provided, the number of specimens measured is given in parentheses. The specimens upon which this paper is based are kept mainly in the Zoological Museum, Moscow State University, Russia (ZMMU, curator Dr. K.G. Mikhailov). More material from Russia and N-Korea were found in the collection of the Zoological Institute, St.-Petersburg, Russia (ZIN, former curator Dr. V.I. Ovtsharenko, recent curator Dr. V.A. Krivokhatskiy), e.g., N-Korean specimens identified by the prominent Russian arachnologist, the late Prof. D.E. Kharitonov. Unfortunately, in some specimens the female genitalia were removed, and the respective slides were not found. Other materials are kept in: IZW Institute of Zoology, Warszawa, Poland (former curator Prof. Dr. J. Prуszyсski); NHMB Natural History Museum Budapest, Hungary (former curator Dr. J. Mahunka, current curator Mr. T. Szuts); JWC Jцrg Wunderlichs (Straubenhardt, Germany) personal collection; OC Dr. T.I. Oligers personal collection (Nizhnesvirskiy Reserve, Leningrad Area, Russia); PC A.V. Ponomarevs personal collection (Razdorskaya Village, Rostov Area, Russia); PSU Perm State University, Perm, Russia (curator Dr. S.L. Esyunin); SZM Siberian Zoological Museum, Novosibirsk, Russia (former curator Dr. D.V. Logunov, current curator Ms. G.N. Azarkina); TNU Taurida National University, Simferopol, Crimea, Ukraine (curator N.M. Kovblyuk); ZMT Zoological Museum, Turku University, Finland (curator M. Saaristo). The following abbreviations for geographical locations are used in the literature data: RUSSIA. ALT Altai Province, AMR Amur Area, ARKh Arkhangelsk Area, AST Astrakhan Area, BLG Belgorod Area, BRN Bryansk Area, BUR Buryatian Republic, ChLB Chelyabinsk Area, ChIT Chita Area, ChVSh Chuvashian Republic, EKB Ekaterinburg (former Sverdlovsk) Area, IRK Irkutsk Area, KBA Kabardino-Balkarian Republic, KhAB Khabarovsk Province, KLM Kalmykian Republic, KLZh Kaluga (Kaluzhskaya) Area, KMCh Kamchatka Area, KMR Kemerovo Area, KOM Komi Republic, KRG Kurgan Area, KRL Karelia (Karelian Republik), KRN Krasnoyarsk Province, KRS Krasnodar Province, KRV Kirov Area, KUR Kursk Area, LNN Leningrad Area, MGD Magadan Area, MRM Murmansk Area, MRS Mariyskaya (Mari-El) Republic, MSK Moscow Area, NVG Novgorod Area, NVS Novosibirsk Area, ORL Orel Area, ORN Orenburg Area, PRI Primorie (=Maritime) Province, PRM Perm Area, PSK Pskov Area, RST Rostov Area, RZN Ryazan Area, SAKh Sakhalin Area (including Kurile Islands), SMR Samara (former Kuybyshev) Area, STV Stavropol Provice, TLS Tula Area, TMB Tambov Area, TMN Tyumen Area, TMS Tomsk Area, TTR Tartar (=Tatar) Republic, TVR Tver (former Kalinin) Area, ULN Ulyanovsk Area, VRN Voronezh Area, YaKT Yakutian (Sakha) Republic. Correct physiographical sequence of the regions given above: MRM, KRL, LNN, PSK, NVG, TVR, MSK, KLZh,

Misidentifications
Clubiona hilaris Simon, 1878 Azerbaijan [Guseinov, Rubtsova, 2001]. The specimen is identified herein as C. alpicola. Clubiona canadensis Emerton, 1890 Cisbaikalia [Izmailova, 1989b s. Mikhailov, 1992a]. Specimen not examined. Clubiona rosserae Locket et Millidge, 1953 Transbaikalia [Izmailova, 1989b s. Mikhailov, 1992a]. Specimen not examined.

Taxonomic part
Genus CLUBIONA Latreille, 1804 Subgenus Paraclubiona Lohmander, 1945 Clubiona corticalis (Walckenaer, 1802)
LITERATURE DATA. AZERBAIJAN [Dunin, Mamedov, 1992]. MISIDENTIFICATIONS. RUSSIA. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993-ld]. These records are clearly outside the known range of this species. UKRAINE: the Crimea: Dzhankoi [Apostolov, Onchurov, 1998; Onchurov, 1998]. DISTRIBUTION. Ukraine (the Carpathians), Caucasus: Azerbaijan, Armenia. The record of this species in the steppe part of the Crimea by Onchurov [1998] is rather doubtful and requires confirmation. S-, C-Europe, S-Sweden.

Subgenus Japoniona Michailov, 1990 Clubiona (Japoniona) japonica L. Koch, 1878
Clubiona flavipes (nec Saito, 1939, misidentified) Mikhailov, 1990b: 144, figs. 1011 ().


The spider genus Clubiona in the fauna of the former USSR: 2003 update
C. japonica Hayashi, Yoshida, 1991: 42, figs. 1214 (). MATERIAL. 1 (NHMW), labelled as: L. Koch coll. Japan. 1884. 825. Steindacher don.

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REMARKS. Earlier [Mikhailov, 1990b], I treated true C. japonica as C flavipes and C. coreana Paik, 1990 as C. japonica. The correct placement for the Russian C. japonica and C. flavipes was proposed by Hayashi & Yoshida [1991]. During a visit to the Naturhistorisches Museum Wien (Austria) in 1995, I was able to examine the C. japonica specimen identified by L. Koch, and the opinion of my Japanese colleagues was confirmed. LITERATURE DATA. RUSSIA. SAKh: Sakhalin [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (islands): SSakhalin, Kunashir. All Japan.

T

HE TRIVIALIS - GROUP

Clubiona (Clubiona) trivialis C.L. Koch, 1843
LITERATURE DATA. RUSSIA. KRL: Kivach Reserve [Tsellarius, 1993]. LNN: Nizhnesvirskiy Reserve [Oliger T., 1995, 1996]. ARKh: Pechora River Delta [Mazura, 2000]. KOM [Mikhailov, 1992b]. ChLB: Troitsky Reserve [Esyunin, Efimik, 1995]. TMN: Yuganskiy Reserve [Mikhailov, 1992b; Esyunin, 1996], Polar Urals [Koponen et al., 1998]. KRN: environs of Mirnoe [Rybalov et al., 2002]. MGD: environs of Magadan [Marusik et al., 1992a; Mikhailov, Marusik, 1995]. SAKh: Sakhalin [Mikhailov, 1992b; Marusik et al., 1993b]. ESTONIA [Vilbaste, 1974, 1993]. LATVIA [Mikhailov, 1992b]. LITHUANIA [Pupiska, 1939; Valenta, Kaarskyte, 1990; Vilkas, 1992-ld; Relys, 1994]. UKRAINE. Polesye [Evtushenko, 1991c, 1993a]. MISIDENTIFICATIONS. KLM [Minoranskiy, Ponomarev, 1984] (examined; see under C. juvenis). DISTRIBUTION. Russia European part (except the extreme South), C- & NE-Siberia, Sakhalin. Baltic countries, Belarus, Ukraine. S-, C- & N-Europe. Japan (Hokkaido).

Clubiona (Japoniona) coreana Paik, 1990
Clubiona coreana Paik, 1990a: 8991, figs. 115 (); C. japonica (nec L. Koch, 1878, misidentified) Mikhailov, 1990b: 143, figs. 16 (), 1992b; Song et al., 1999: 416, figs. 246J K (), 249IJ (); C. coreana Song et al., 1991: 6768, fig. 2AD (); Song et al., 1999: 415, figs. 245BC (), 247KL (). MATERIAL. RUSSIA, Maritime Prov.: 12 , 11 (OC, ZMMU, SZM), Lazovskiy Distr. and Reserve, 19751982, TO; 1 (ZMT), Ussuri Reserve, 43њ39N, 132њ33E, 2931.07.1998, leg. SK, det. YM. More specimens were collected in KhAB: Bolshekhekhtsyrskiy Reserve and PRI: environs of Anisimovka Village, Khasan Distr., and Sikhote-Alinsky Reserve.

Clubiona (Clubiona) amurensis Michailov, 1990
LITERATURE DATA. RUSSIA. PRI [Mikhailov, 1992b]. REMARKS. The male of C. amurensis from Hokkaido figured by Matsuda [1991] is not conspecific with true C. amurensis. A specimen from Kunashir (see below) belongs to this undescribed Japanese species. DISTRIBUTION. Russia S-Far East: Khinganskiy Reserve.
Clubiona amurensis Mikhailov, 1990b: 148, figs. 2122 ();

LITERATURE DATA. RUSSIA. PRI [Mikhailov, 1992b; Kim, Kurenshchikov, 1995-ld]. REMARKS. See Remarks under the former species. The correct placement of Russian C. japonica was indicated by Paik [1990b] and Hayashi & Yoshida [1991]. Their opinion is accepted here (see above). DISTRIBUTION. Russia S-Far East (continental). S-Korea, China (Liaoning).

Clubiona vigil Karsch, 1879
Clubiona vigil Mikhailov, 1990b: figs. 79 (); Hayashi, Yoshida: 42, figs. 911 ().

Clubiona (Clubiona) cf. amurensis Michailov, 1990
Clubiona amurensis Matsuda, 1991: 2224, figs. 17 (). MATERIAL. RUSSIA: S-Kuriles: 1 (ZMMU), Kunashir Isl., S part, 2.5 km N of Golovnino, 43њ46.01N, 145њ32.02E, 01 & 21.09.1997, YM.

REMARKS. According to Hayashi & Yoshida [1991], specimens from Japan and Russia belong to different species. Examination of the type species is necessary. LITERATURE DATA. RUSSIA. SAKh: Kunashir Isl. [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia Kuriles: Kunashir Isl. All Japan, S-Korea, China (?).

REMARKS. See Remarks under the former species. DISTRIBUTION. Russia S-Kuriles: Kunashir Isl. Japan: Hokkaido.

Clubiona (Clubiona) basarukini Michailov, 1990
Clubiona basarukini Mikhailov, 1990b: 146147, figs. 1216 (). C. basarukini Matsuda, 1991: 2426, figs. 811 ().

Subgenus Bucliona Benoit, 1977 (=Bicluona Michailov, 1994) Clubiona (Bucliona) jucunda (Karsch, 1879)
Clubiona jucunda Paik, 1990b: 7173, figs. 2839 (); Mikhailov, 1994: 5456, figs 1(13), 2 (13) (); Song et al., 1999: 425, figs. 246P (), 249M ().

LITERATURE DATA. RUSSIA. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b-ld]. DISTRIBUTION. fUSSR Russia: Transbaikalia, SSakhalin. N-Mongolia: Selenga Aimak. Japan: Hokkaido.

LITERATURE DATA.RUSSIA. PRI: Lazovski Reserve [Mikhailov, 1994]. SAKh: Kunashir [Mikhailov, 1992b, 1994; Marusik et al., 1993b-ld]. DISTRIBUTION. Russia: S-Far East (continental & island). Japan, N- & S-Korea, China (widely [Song et al., 1999]).

Clubiona (Clubiona) diversa O. Pickard-Cambridge, 1862
MATERIAL. KAZAKHSTAN: 1 (SZM), Pavlodar Area, Bayan-Aulskiy Distr., environs of Kyzyl-Tau Village, meadow, 11.06.1991, OL.


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UKRAINE: 1 , 2 (ZMMU), The Crimea, btw. Saki & Yevpatoria, 45њ09.37N, 33њ30.044E, 18.03.2002, leg. et det. YM.

LITERATURE DATA. RUSSIA. LNN [Mikhailov, 1992b]: Nizhnesvirskiy Reserve [Oliger T., 1995, 1996]. KRS: Caucasian Reserve [Mikhailov, 1992b]. BShK [Efimik, 1997]: Bashkirian Reserve [Efimik, 1995a; Esyunin, Efimik, 1995]. ChLB: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. NVS [Mikhailov, 1992b]. KRN: Khakassia [Mikhailov, 1992b]. Tuva [Mikhailov, 1992b; Logunov et al., 1998; Marusik et al., 2000]. MGD: environs of Magadan [Marusik et al., 1992-ld]; Upper Kolyma [Marusiket al., 1992;Marusik, 1994; Mikhailov, Marusik, 1995]. Transbaikalia [Danilov, 1999-ld]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. KhAB: Okhotsk Distr. [Marusik et al., 1992; Mikhailov, Marusik, 1995]. SAKh: Sakhalin [Marusik et al., 1993b]; Middle Kuriles [Kamura, 2001]. LITHUANIA [Vilkas, 1992-ld; Relys, 1994-ld; Relys, Dapkus, 2001; Relys et al., 2002]. UKRAINE. ZAK [Mikhailov, 1992b]. KAZAKHSTAN. Akmolinsk, Alma-Ata, East Kazakhstan areas [Mikhailov, 1992b]; Saur Mt. Ridge [Eskov, Marusik, 1995]. DISTRIBUTION. Russia European part, Caucasus, Siberia, Kuriles. Estonia, Lithuania, Belarus, Ukraine, Azerbaijan, Kazakhstan. S-, C- & N-Europe, Mongolia, S-Korea, Japan.

More specimens were collected in Tuva, ChIT (Kyra Town), and Sakhalin.

LITERATURE DATA. LNN: Nizhnesvirskiy Reserve [Oliger T., 1996]. KUR: Tsentralno-Chernozyomniy Reserve [Polchaninova, 2001]. SMR [Krasnobaev, Matveev, 1993]: Zhiguli Reserve [Krasnobaev, 1996]. PRM [Esyunin, Efimik, 1995]. ChLB: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. NVS [Mikhailov, 1992b]. Tuva [Mikhailov, 1992b]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1997]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. KhAB: S part [Kim, Kurenshchikov, 1995]. SAKh: Sakhalin [Marusik et al., 1993b]. UKRAINE. Askania-Nova Reserve [Polchaninova, 1995]; Chernomorskiy Reserve [Polchaninova, 1997]; Polesye [Evtushenko, 1991b,c, 1993b]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996-ld]. DISTRIBUTION. Russia C- & N-European part, W& S-Siberia. Estonia, Belarus, Ukraine, Kirghizia. S-, C- & N-Europe, Mongolia: Selenga Aimak.

Clubiona (Clubiona) transbaicalica Michailov, 1992
Clubiona (Clubiona) transbaicalica Mikhailov, 1992b: 67, fig. 1 ().

Clubiona (Clubiona) juvenis Simon, 1878
LITERATURE DATA. LITHUANIA [Mikhailov, 1992b]. UKRAINE: DNTs [Prokopenko, 2000], Berdyansk [Mikhailov, 1992b]. DISTRIBUTION. Russia S-European part. Estonia, Lithuania, Ukraine, Uzbekistan. S- & C-Europe, China: Gansu.

LITERATURE DATA. RUSSIA. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]. DISTRIBUTION. Russia Transbaikalia. This species is known only from the type localities.

T

HE MARMORATA - GROUP

Clubiona (Clubiona) marmorata L. Koch, 1866
LITERATURE DATA. LITHUANIA [Pupiska, 1939; Vilkas, 1992-ld; Relys, 1994-ld, 1995]. MOLDAVIA [Mikhailov, 1992b]. MISIDENTIFICATION. RUSSIA. KRS [Sharov et al., 1984]. DISTRIBUTION. Lithuania, Belarus, Ukraine, Moldavia. Not found in Russia. S- & C-Europe.

Clubiona (Clubiona) rostrata Paik, 1985
Clubiona rostrata Paik, 1985: 35, figs. 1018 (); C. maikoae Hayashi, 1985: 3639, figs. 18 (); C. rostrata Hayashi, 1987: fig. 14 (, = C. maikoae); Mikhailov, 1990b: figs. 1720 (); Paik, 1990b: 7576, figs. 4852 (); Song et al., 1999: 427, figs. 251IJ (), 253JK (). MATERIAL. RUSSIA, Maritime Prov.: 3 (ZMMU), Khasan Distr., environs of Andreevka Village, 42њ3536N, 131њ13E, 1115.08.1998, leg. et det. YM; 3 , 7 (OC, ZMMU), Lazovskiy Distr., 19751982, 1984, TO; 1 (ZMMU), Lazovskiy Reserve, Korpad Camp, 43њ16N, 134њ08E, 0609.08.1998, SK. DOUBTFUL LOCALITY. RUSSIA: 1 (PC), Belgorod Area, N suburb of Belgorod, potato field, pitfall traps, 27.08.1977, A.V. Prisnyi. More specimens were collected in KhAB: Bolshekhekhtsirskiy Reserve and PRI: environs of Anisimovka.

Clubiona (Clubiona) hyrcanica Michailov, 1990 Figs. 12.
Clubiona hyrcanica Mikhailov, 1990a: 303304, figs 58 (). MATERIAL. 1 1 (ZMMU), SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001, YM.

REMARKS. The record from Belgorod Area is extremely far from the main range of C. rostrata. The specimen identification seems to be correct, therefore I suspect that the label locality data is wrong. LITERATURE DATA. RUSSIA. KhAB: Nanayskiy Distr. [Mikhailov, 1992b; Kim, Kurenshchikov, 1995-ld]; Bolshoi Khekhtsir Mt. Range [Kim, Kurenshchikov, 1995-ld]. PRI [Mikhailov, 1992b]. DISTRIBUTION. Russia: S-Far East (continental & island). Japan, N- & S-Korea, China: Zhejiang, Liaoning, Jilin, Heilongjiang.

Clubiona (Clubiona) subtilis L. Koch, 1867
MATERIAL. RUSSIA: 1 (ZMMU), Krasnodar Province, 10 km NW of Starokorsunskaya, edge of wheat field, 28.04.2000, R. Seifulina.

DIAGNOSIS(female). Previously, this species was known only from the male. The female, descrbed here for the first time, differs from the closely related species C. marmorata, in the absence of the central paired epigynal grooves, the nonsclerotized posterior lateral angles of the epigyne and in the different general orientation of the copulatory tubes. DESCRIPTION. Female. Total length 5.85. Carapace 2.55 long, 1.90 wide, ratio 1.34. Carapace and legs strawcoloured. Leg measurements: F Pt Ti Mt T I 1.30 0.80 1.00 0.75 0.45 II 1.50 0.88 1.18 0.75 0.50 III 1.25 0.68 0.75 1.00 0.43 IV 1.75 0.88 1.38 1.68 0.58 Leg armature: Ti II v.1.2, III d.2, v.1, Mt I v.0, III d.2.0.2, lat.1.2, v.2.2, IV d.2.1.2, lat.2.2, v.2.2. Abdomen pale brown, 3.20 long, 1.85 wide, ratio 1.73. Epigyne as in Figs. 12.


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UKRAINE. Polesye [Evtushenko, 1993b]. MOLDAVIA [Esyuninet al., 1993, 1995; Penev et al., 1994]. DISTRIBUTION. Russia N-Caucasus. Baltic countries, Belarus, Ukraine, Moldavia. S-, C- & N-Europe, Algeria.

The genevensis-subgroup Clubiona (Clubiona) genevensis L. Koch, 1866
MATERIAL. UKRAINE: 5 , 2 (ZMMU, SZM, TNU), The Crimea, Sakskiy Distr., Pribrezhnaya Railway Station, dry meadow & haloratophytum, 28.05 24.06, 21.0905.11, 0318.12.2000, NK. AZERBAIJAN: 1 (ZMMU), Apsheron Peninsula, env. of Gyurgyan Vill., 40њ24N, 50њ16E, semidesert, 17.04.2001, YM; 1 (ZMMU), Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001, YM. KAZAKHSTAN: 1 (ZMMU), Pavlodar Area, Bayan-Aulskiy Distr., Babaly Mts., stony debris, 27.08.1990, OL.

REMARKS. As I have previously indicated [Mikhailov, 1992a], the available material does not allow me to distinguish between the two very close species C. genevensis and C. vegeta SiFigs. 12. Clubiona hyrcanica, genitalia: 1 epigyne, 2 vulva. mon, 1878. Therefore, all the records of C. vegeta Рис. 12. Clubiona hyrcanica, гениталии : 1 эпигина, 2 вульва. from the fUSSR are here ascribed to C. genevensis. LITERATURE DATA.AZERBAIJAN: Lenkoran [GuseiLITERATURE DATA. RUSSIA. BShK: Bashkirian Renov, 1998, 1999]. serve [Mikhailov, 1992b]. KRN: Khakassia [Mikhailov, 1992b]. DISTRIBUTION. Azerbaijan: Lenkoran, Hyrcan. AZERBAIJAN: Apsheron Peninsula, Lenkoran [Guseinov, 1998, 1999]. KAZAKHSTAN. Saur Mt. Ridge [Mikhailov, 1992b; T HE BREVIPES - GROUP Eskov, Marusik, 1995]. KIRGHIZIA [Zonshtein et al., 1996-ld: C. genevensis + Clubiona (Clubiona) brevipes Blackwall, 1841 C. vegeta]. DISTRIBUTION. Russia S-European part, S-Urals, MATERIAL. UKRAINE: 1 (ZMMU), the Crimea, Krimskiy S-Siberia (Khakassia). Latvia, Ukraine, Azerbaijan, ArmeState Natural Reserve, Asport Camp, broadleaved forest, 28.06. nia, N-, W- & E-Kazakhstan, Turkmenistan, Kirghizia, Tad2001, NK. jikistan. GEORGIA: 1 (ZIN), Kolkhida, Bdzho-Bdzho, 15.06.1969, E. Popov. S-, C- & N-Europe, Morocco, Algeria, Palestine. LITERATURE DATA. RUSSIA. STV [Mikhailov, 1992b]. MISIDENTIFICATION. MRS [Matveev, 1991; KrasTHE LUTESCENS - GROUP nobaev, Matveev, 1983-ld]. This record is clearly outside the known range of this species. Clubiona (Clubiona) lutescens Westring, 1851 DISTRIBUTION. Russia S-European part, Ukraine (The Crimea), Georgia, Azerbaijan. MATERIAL. RUSSIA: 5 , 4 (PC), Belgorod Area, S-, C- & N-Europe. New to the fauna of Ukraine.

Clubiona (Clubiona) alexeevi Michailov, 1990
LITERATURE DATA. RUSSIA. KBA [Mikhailov, 1992b]. DISTRIBUTION. Russia: N-Caucasus: C part.

T

HE COMTA - GROUP

Clubiona (Clubiona) comta C.L. Koch, 1839
REMARKS. As has been indicated in several catalogues [Prуszyсski, Starкga, 1971, etc.], the spelling C. comta is correct, whereas C. compta is an invalid emendation. LITERATURE DATA. RUSSIA. KRS [Mikhailov, 1992b]. ESTONIA [Vilbaste, 1974: C. compta; 1993: C. compta]. LITHUANIA [Pupiska, 1939: C. compta; Vilkas, 1992ld: C. compta].

environs of Kustovoe and Borisovka, 1986, 19961997, leg. et det. AP; 1 (SZM), Tyumen Area, Zavodouksk Distr., Lybaevo Vill., 06.1988, DL; 1 (ZMMU), Tomsk Area, 25 km of Kargasok Village, deciduous forest, under bark, 12.08.1983, AZ. GEORGIA: 1 (ZMMU), 40 km W of Mestia, Kherkhvashi E of Narka (=Naki), 12501700 m, Quercus, Fagus, Carpinus, Picea, Abies etc. forest, litter & bark, 21.0821.09.1986, SG; 1 , 4 (ZMMU, SZM), Algeti Reserve W of Manglisi, Fagus, Picea, Acer, etc. forest, 14001450 m, litter & under bark, 16 18.05.1987, SG & KE. KAZAKHSTAN: Pavlodar Area: 1 (ZMMU), environs of Zhelezenka Village, Irtysh River floodplain, Populus-Salix forest with blackberry, 06.08.1988, OL; 2 (ZMMU), Uspenskiy Distr., 15 km E of Bogatyr Sovkhoz, Betula-Populus tremula groves, 19.07.1990, OL. IRAN: 2 , 4 (ZMMU), Mazandaran Prov., N of Jahaver-Deh Vill., 36.872њN, 50.467њE, 09.06.2000, YM. More specimens were collected in Russia: Altais, Tuva, and Kazakhstan: VKO.


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GEORGIA[Mkheidze, 1997:C. frutetorum & C. liachviana]. AZERBAIJAN. Apsheron Peninsula [Guseinov, 1998], Lenkoran [Guseinov, 1998, 1999]. TURKMENISTAN. SW Kopetdagh Mts. [Mikhailov, Fet, 1994]. DISTRIBUTION. Ukraine (Carpathians, the Crimea),Georgia, Azerbaijan, Armenia, Turkmenistan (Kopetdagh Mts.). C-Europe, Bulgaria, Romania.

LITERATURE DATA. RUSSIA. KRL: Kivach Reserve [Tsellarius, 1993]. LNN: Nizhnesvirskiy Reserve [Oliger T., 1995, 1996]. TVR: Tsentralno-Lesnoi Reserve [Zheltukhina, 1995]. MSK [Seifulina, Chernyshev, 2001]. TLS [Esyunin et al., 1993, 1995; Penev et al., 1994; Tanasevitch, 1999]. RZN [Mikhailov, 1992b]. KUR: Tsentralno-Chernozyomniy Reserve [Polchaninova, 2001]. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993]. ChVSh [Sidorenko, Dunin, 1997]. ULN [Krasnobaev, 1999]. TTR: Volzhsko-Kamskiy Reserve [Esyunin et al., 1993, 1995]. SMR [Krasnobaev, Matveev, 1993, 1999]: Zhiguli Reserve [Krasnobaev, 1996]. KOM [Mikhailov, 1992b]. PRM [Mikhailov, 1992b; Pakhorukov et al., 1995; Esyunin, Efimik, 1996]. EKT: Visimskiy Reserve [Esyunin, Ukhova, 2001]. BShK [Efimik, 1997]: Shulgan-Tash Reserve [Efimik, Gulyashchikh, 1995]. ChLB [Efimik, Zolotarev, 1998]: Troitskiy Reserve [Esyunin, Pakhorukov, 1992], Ilmenskiy Reserve [Polyanin, Lagunov, 1992]. ORN [Esyunin, Efimik, 1996; Efimik et al., 1997]. KRG [Mikhailov, 1992b]. TMN: Yarkovo Distr. [Mikhailov, 1992b], Yuganskiy Reserve [Esyunin, 1996]. NVS [Mikhailov, 1992b]. KMR [Mikhailov, 1992b; Romanenko, 1998]. ALT [Marusik et al., 1996*; Azarkina, 1999]. KRN: environs of Mirnoe [Marusik et al., 2002b; Rybalov et al., 2002]; Khakassia [Mikhailov, 1992b]. Tuva [Logunov et al., 1998: ?C.l.; Marusik et al., 2000]. YaKT: Sangar [Mikhailov, 1992b; Marusik et al., 1993a-ld]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]. KhAB: Amur River between Nikolaevsk and Komsomolsk, Bikin Distr. [Kim, Kurenshchikov, 1995]; Ulchskiy Distr. [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b-ld]. ESTONIA [Vilbaste, 1974, 1993]. LITHUANIA [Pupiska, 1939; Mikhailov, 1992b; Vilkas, 1992-ld; Relys, 1994-ld]. BELARUS. Pripyatskiy National Park [Zhukovets, 1997]. UKRAINE. DNTs: Khomutovskaya Steppe Reserve [Polchaninova, 1993]; Luganskiy Reserve [Prokopenko, 1999a]; Polesye [Evtushenko, 1991a,b, 1993a,b]; Askania-Nova Reserve [Polchaninova, 1995]. ZhTM: Polesskiy Reserve [Guryanova, 1994]. GEORGIA [Mkheidze, 1997]. AZERBAIJAN. Lenkoran [Guseinov, 1998, 1999]. KAZAKHSTAN. Saur Mt. Ridge [Mikhailov, 1992b; Eskov, Marusik, 1995]. DISTRIBUTION. Russia European part, Caucasus, Urals, Siberia, S-Far East (continental & island). Baltic countries, Belarus, Ukraine, Moldavia, Georgia, Azerbaijan, Armenia, N- & E-Kazakhstan. S-, C- & N-Europe, Japan, N-America (Pacific coast). New to the fauna of Iran.

Clubiona (Clubiona) frutetorum L. Koch, 1867
MATERIAL. RUSSIA: 1 (PC), Belgorod Area, 8 km SW of Borisovka, environs of Krasivo Village, pine forest, 17.06.1997, leg. et det. AP; 1 (ZMMU), Orenburg Area, Sol-Iletsk Distr., Shybyndy Ravine, 22.08.2001, S.L. Esyunin. UKRAINE: 1 (ZMMU), the Crimea, Krymskiy State Natural Reserve, Tarier Kordon locality, 30.06.2001, NK. KAZAKHSTAN: 1 (ZMMU), Semipalatinsk Area, Ayaguz Distr., 16 km W of Kalgutty, Betula forest, 13.10.1991, A.A. Fedorov, T.V. Iskakova, A.A. Zyuzin.

LITERATURE DATA. RUSSIA. KRL: Kivach Reserve [Tsellarius, 1993]. LNN [Mikhailov, 1992b]: Nizhnesvirskiy Reserve [Oliger T., 1996]. KUR: Tsentralno-Chernozyomniy Reserve [Polchaninova, 2001]. MRS [Mikhailov, 1992b]. SMR [Krasnobaev, Matveev, 1993]. KOM [Mikhailov, 1992b]. ChLB: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. ESTONIA [Vilbaste, 1974]. LITHUANIA [Pupiska, 1939; Vilkas, 1992-ld; Relys, 1994-ld]. UKRAINE. ChRK: Kanevskiy Reserve [Mikhailov, 1992b]; Polesye [Evtushenko, 1991b, 1993a,b]; KhRS: Chernomorskiy Reserve [Polchaninova, 1998]; DNTs: Luganskiy Reserve [Prokopenko, 1999a], Donetsk [Prokopenko, 1999b]. DISTRIBUTION. Russia European part, S-Siberia. Baltic countries, Belarus, Ukraine, Moldavia, Kazakhstan. All Europe.

Clubiona (Clubiona) golovatchi Michailov, 1990 Figs. 3, 10, 14.
Clubiona golovatchi Mikhailov, 1990a: 307310, figs 1315 (, in part, ). MATERIAL. RUSSIA: 1 (holotype, ZMMU Ta-4439), Krasnodar Prov., Sochi, Lazarevskoe, 28.10.1981, SG; 1 (paratype, ZMMU), Stavropol Prov., W of Zheleznovodsk, 29.05.1982, SG; North Osetia: 2 (paratypes, ZMMU), Tsey Mt. Ridge, 6.0621.07, 21.078.08.1985, S.K. Alekseev; 1 (paratype, ZMMU), S of Ordzhonikidze, btw. Chmi & Baltik, 2.06.1982, SG; 1 (paratype, ZMMU), Daghestan, Samur Valley, Garakh, ca. 35 km SW Magaramkent, 23.10.1987, SG. AZERBAIJAN: 1 (paratype, ZMMU), Chilisa ca. 7 km N of Kelbajar, 31.05.1987, SG & KE. ARMENIA: 1 (paratype, ZMMU), Kafan Distr., near Kadjaran, Megri Mt. Ridge, 27.04.1983, SG.

Clubiona (Clubiona) alpicola Kulczyсski, 1881
Clubiona liachviana Mkheidze, 1997: 175176, figs. 345, 346 (), syn.n. C. frutetorum Mkheidze, 1997: 176, fig. 347 (). MATERIAL: GEORGIA: 3 (ZIN), Adigeni Distr., Benara Vill., right bank of Abastumani River, 06.06.1949, E. Kiryanova. More specimens were collected in Azerbaijan.

REMARKS. Judging from the original drawings, C. liachviana Mkheidze, 1997 decribed from Gori and Bagdadi, Georgia, is a junior synonym of C. alpicola. As has been suggested previously [Mikhailov, 1990a, 1992b], all records of C. frutetorum from the Caucasus and Kopetdagh Mts. should be ascribed to C. alpicola. LITERATURE DATA. UKRAINE. The Crimea [Mikhailov, 1992b].

DIAGNOSIS. The male is close to C. saxatilis L. Koch, 1867 (= C. dvoraki Miller, 1943) and C. alpicola, but differs from the former species in the longer embolus and from the latter in the presence of the rib dividing the sclerotized and membranous parts of the embolus. In C. alpicola, the rib divides two equally or subequally sclerotized parts of the embolus. For the diagnosis of the female, see Mikhailov [1990a]. REMARKS. The type series of C. golovatchi turned out to include males of two species; the new one is decribed below as C. mazandaranica. All females in the type series belong to the former species.


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289

3

4

5

6

7

8

9

Figs. 39. Clubiona spp., left (3, 69) & right (45) palp: 3 C. golovatchi, paratype (N Osetia, S of Ordzhonikidze), 45 C. mazandaranica sp.n., paratype (Azerbaijan, Hyrcan Reserve), 67 C. mykolai sp.n., holotype, 89 C. saxatilis (Austria, Innsbruck). 3, 4, 6, 8 ventrally, 5, 7, 9 retrolaterally. Рис. 39. Clubiona spp., левый (3, 69) и правый (45) пальпус: 3 C. golovatchi, паратип (Сев. Осетия, южнее Орджоникидзе), 45 C. mazandaranica sp.n., паратип (Азербайджан, Гирканский заповедник), 67 C. mykolai sp.n., голотип, 89 C. saxatilis (Австрия, Инсбрук). 3, 4, 6, 8 вентрально, 5, 7, 9 ретролатерально.


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K.G. Mikhailov

12 10 11 13

Figs. 1013. Clubiona spp., left (10, 1213) & right (11) palp, dorsally: 10 C. golovatchi, paratype (N Osetia, S of Ordzhonikidze), 11 C. mazandaranica sp.n., paratype (Azerbaijan, Hyrcan Reserve), 12 C. mykolai sp.n., holotype, 13 C. saxatilis (Austria, Innsbruck). Рис. 1013. Clubiona spp., левый (10, 1213) и правый (11) пальпус, дорсально: 10 C. golovatchi, паратип (Сев. Осетия, южнее Орджоникидзе), 11 C. mazandaranica sp.n., паратип (Азербайджан, Гирканский заповедник), 12 C. mykolai sp.n., голотип, 13 C. saxatilis (Австрия, Инсбрук).

LITERATURE DATA. RUSSIA. KRS: environs of Krasnodar [Seifulina, 2002]. DISTRIBUTION. Russia foothills and mountains of the Caucasus Major. Georgia, Azerbaijan, Armenia.

Clubiona (Clubiona) mazandaranica Michailov, sp.n. Figs. 45,11, 15, 1819.
Clubiona golovatchi (partim) Mikhailov, 1990a: figs. 912 (, not !). MATERIAL. Holotype (ZMMU Ta-6594), IRAN, Mazandaran Prov., env. of Javaher-Deh Vill., 36.872њN, 50.467њE, 09.06.2000, YM. Paratypes: AZERBAIJAN: 1 (ZMMU Ta-4863, paratype of C. golovatchi), Istisu ca. 8 km SW of Masally, Quercus, Acer, Carpinus etc. forest, 80140 m a.s.l., litter, under bark & stones, 1920.10.1983, SG; 1 (ZMMU Ta-4864, paratype of C. golovatchi), Hyrcan Reserve, near bridge on Bashara River, Urtica, Sambucus, etc., 28.06.1983, DL; 1 (JWC), IRAN, Mazandaran Prov., env. of Khorran-abad Vill., 36њ43 N, 50њ48E, 0810.06.2000, YM.

DIAGNOSIS. The male of C. mazandaranica sp.n. differs from C. golovatchi in the ridge on the embolus, the uniformly sclerotized embolus, the presence of a protruding region of the tegulum, the wide ventral branch of the TA and the presence of the poorly visible indentation of the dorsal TA branch. The female differs by having wider copulatory tubes and comparatively small atria, which slightly resemble a variation of C. golovatchi (cp. Mikhailov, 1990a: fig. 15). DESCRIPTION. MALE (measurements of paratypes are given in parentheses). Body length 6.05 (6.45 for ZMMU Ta-

4863). Carapace 2.85 (2.632.85) long, 1.88 (1.682.05) wide, ratio 1.52 (1.391.58). Carapace, chelicerae and legs straw-coloured, or chelicerae a little reddish. Palp as in Figs. 45, 11, 15. Lengths of F, Pt, Ti, and Cb are 0.85 (0.780.80), 0.38 (0.380.40), 0.30 (0.300.33), 1.05 (1.051.10), respectively. Leg measurements: F Pt Ti Mt T I 2.30 1.10 2.13 1.58 0.90 (2.13) (1.081.13)(2.032.15)(1.501.58)(0.850.90) II 2.38 1.05 2.25 1.68 0.95 (2.25) (1.081.13)(2.102.25)(1.551.60) (0.93) III 2.00 0.93 1.50 1.80 0.75 (1.931.98) (0.880.93)(1.431.50)(1.701.80)(0.630.70) IV 2.70 1.08 2.28 2.90 0.90 (2.552.60) (1.001.03)(2.052.28)(2.703.00) (0.88) Leg armature: F I d.1.1.3 (rarely 1.1.2), Ti III d.2.2., v.1.1.1 (rarely 1.1). Abdomen pale brown or greyish-brown, 3.25(3.353.75) long, 1.58(1.751.78) wide, ratio 2.06(1.912.11). FEMALE. Body length 7.2. Carapace straw-coloured, 2.78 long, 2.00 wide, ratio 1.39. Chelicerae pale brown. Legs straw-coloured. Leg measurements: F Pt Ti Mt T I 1.88 1.00 1.55 1.13 0.70 II 1.93 1.00 1.63 1.18 0.75 III 1.75 0.88 1.25 1.50 0.60 IV 2.48 1.10 1.98 2.50 0.83 Leg armature: Ti III d.2.2, v.1.1.1 (1.1.0), Mt III d.2.1.2, lat.2.2 (1.2), v.2.2.


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291

14

15

16

16a

17

Figs. 1417. Clubiona spp., left (14, 1617) & right (15) palp, tibial apophysis: 14 C. golovatchi, paratype (N Osetia, S of Ordzhonikidze), 15 C. mazandaranica sp.n., paratype (Azerbaijan, Hyrcan Reserve), 1616a C. mykolai sp.n., holotype, 17 C. saxatilis (Austria, Innsbruck). Рис. 1417. Clubiona spp., левый (14, 1617) и правый (15) пальпус, вырост голени: 14 C. golovatchi, паратип (Сев. Осетия, южнее Орджоникидзе), 15 C. mazandaranica sp.n., паратип (Азербайджан, Гирканский заповедник), 1616a C. mykolai sp.n., голотип, 17 C. saxatilis (Австрия, Инсбрук).

18

19

Figs. 1819. Clubiona mazandaranica sp.n., paratype (Iran, Mazandaran Prov.), genitalia, epigyne (18), vulva (19). Рис. 1819. Clubiona mazandaranica sp.n., паратип (Иран, пров. Мазандаран), гениталии , эпигина (18), вульва (19).

Abdomen pale brown, 4.38 long, 2.38 wide, ratio 1.84. Epigyne as in Figs. 1819. LITERATURE DATA. AZERBAIJAN: Lenkoran [Guseinov, 1998, 1999: both as C. golovatchi]. DISTRIBUTION. Azerbaijan: Lenkoran Prov. Iran: Mazandaran Prov. DERIVATIO NOMINIS. The species is named after the type locality, Mazandaran Prov. of Iran.

Clubiona (Clubiona) mykolai Michailov, sp.n. Figs. 67, 12, 16, 16a, 2022.
MATERIAL. Holotype (ZMMU Ta-6595), UKRAINE, the Crimea, environs of Simferopol Town, near Kirpichnoe Village, windbreak, 1431.05.1997, NK. Paratypes: UKRAINE, The Crimea: 1 (ZMMU), environs of Bakhchisarai, Skalistoe Village, 01.11.1995; 1 (ZMMU), Simf-


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20

21

22
eropol Distr., 1.52 km N of Fermanovo Vill., edge of Quercus forest, 1626.07.2000; Krimskiy (Krymskiy) State Natural Reserve: 1 (TNU), Kordon Bukovskogo, Fagus forest, ca. 1100 m a.s.l., 12.06.2000, 2 (ZMMU), Kordon Asport, mixed broadleaved forest, 28.06.2001, 1 (TNU), same locality, bank of Alma River, 21.08.2001; 1 (ZMMU), Nikitskaya Yaila Mts., ca. 900 m a.s.l., Pinus, Quercus, Carpinus, Acer forest, 1331.03.2001; 1 (ZMMU), Yalta, Massandra Park, Bambusa sp. near stream, 08 15.07.2000; all leg. by NK. COMPARATIVE MATERIAL. C. saxatilis (labelled as C. dvoraki) (Figs. 89, 13, 17): 1 , 1 (coll. K. Thaler), Insbruck, Hallthal, 1965 (cp. Thaler, 1981: 119).

Figs. 2022. Clubiona mykolai sp.n., paratypes, genitalia (2021 Ukraine, Crimea, Krimskiy Reserve, Asport Camp), epigyne (20, 22), vulva (21). Рис. 2022. Clubiona mykolai sp.n., паратипы, гениталии (2021 Украина, Крым, Крымский заповедник, кордон Аспорт), эпигина (20, 22), вульва (21).

DIAGNOSIS. The male of C. mykolai sp.n. is most close to the Central-European C. saxatilis L. Koch, 1867 (cp. Figs. 6 and 8), but differs in the following characters: the presence of a small keel at the upper margin of the embolus (such that the keel is more pronounced in C. golovatchi Michailov, 1990: see Fig. 3); the bulb is narrower, and the ventral TA branch is flatter, thinner and transparent. The female of C. mykolai sp.n. is most close to the West-European C. hilaris Simon, 1878 and differs in the following characters: the longer, thinner, more sinuate 2nd spermathecae; the atrium and 1st spermathecae are more distant from each other, being connected with a thin tube; the less rounded 1st spermathecae (seen in dorsal view). REMARKS. A more detailed comparison of C. mykolai sp.n. with C. saxatilis, C. golovatchi / C. mazandaranica and C. hilaris seems to be necessary.

1. The TA in the new species is thinner and more transparent than in C. saxatilis and C. hilaris, as figured by K. Thaler [1981: Abb. 33 & 34; see also Figs. 16, 16a, 17 here]. 2. In several characters, C. mykolai sp.n. appears to be closer to C. golovatchi / C. mazandaranica. The main differences of the new species are: the less pronounced keel on the embolus (see Fig. 6; missing in C. saxatilis and C. golovatchi, but present in C. mazandaranica) and the indentation of the TA dorsal branch (missing in C. saxatilis and C. golovatchi, but present in C. mazandaranica). In C. golovatchi, the loops of the seminal duct are more clearly visible at the distal tip of the tegulum. Median tubercle at the tegular tip is present in C. mykolai, C. golovatchi and C. mazandaranica (see Figs. 3, 4 and 6; almost missing in C. saxatilis), but is more heavily sclerotized in C. golovatchi. 3. A rib subdividing sclerotized and membranous parts of the embolus present in C. saxatilis, C. golovatchi and C. frutetorum L. Koch, 1867. In C. saxatilis, the rib crosses the embolus at a point approximately one third of its length from the proximal end, in C. golovatchi it crosses the point approximately equal to half of its length. 4. In C. saxatilis and C. golovatchi, the TA ventral branch is slightly shorter than the dorsal one, and vice versa in the new species. The TA dorsal branch is wider in C. saxatilis and, more so in C. mazandaranica, than it is in C. mykolai sp.n. The characters discussed above are summarized in the following table:


The spider genus Clubiona in the fauna of the former USSR: 2003 update
Char acter The keel at the upper em bolic mar gin Median tubercle of the apical par t of tegulum The r ib dividing s cler otized and m embr anous parts of the em bolus The TA ventr al branch

293

C. mykolai
pr esent ( Fig. 6) pr esent ( Fig. 6)

C. hilaris
absent present

C. saxatilis
absent alm os t absent ( Fig. 8 and, e specially, 9) pres ent, at f irst 1/3 of embolus length (Fig. 8) wider than in C. dvoraki ; s hor ter than dor sal branch thicker, les s transparent without indentation

C. golovatchi
absent present pr esent, at half of em bolus length ( Fig. 3) shor ter than dor sal branch thicker, les s transparent without indentation

C. maz andaranica
pres ent, m or e pronounced ( Fig. 4) pr esent, well-pronounced (F igs 45) absent wider than in C. dvoraki ; subequal in length with dor sal branch thicker, less transparent with s mall indentation (F ig. 11)

absent

absent subequal in length with dor sal branch thicker, les s transparent without indentation

longer than dor sal branch thin, transparent with indentation (F igs 12, 16, 16a)

The TA ventr al branch The TA dor sal branch

DESCRIPTION. MALE. Body length 5.50. Carapace 2.35 long, 1.90 wide, ratio 1.24. Carapace and chelicerae reddish. Chelicerae 1.05 long. Palp as in Figs. 67, 12, 16, 16a. Lengths of F, Pt, Ti, and Cb are 0.63, 0.35, 0.28, and 0.85, respectively. Legs straw-coloured. Leg measurements: F Pt Ti Mt T I 1.73 1.73 1.65 2.13 0.65 II 0.93 0.95 0.75 0.90 0.65 III 1.53 1.55 1.05 1.70 0.50 IV 0.65 0.65 0.50 0.68 0.68 Leg armature: Mt III d.2.1.2, lat.2.2 (1.2), v.2.2. Abdomen pale brown, 3.10 long, 1.80 wide, ratio 1.72. FEMALE (8). Body length 6.09+0.62. Carapace 2.65+0.18 long, 1.89+0.17 wide, ratio 1.40+0.08. Carapace and legs straw-coloured, in several specimens carapace a little bit darker. Chelicerae pale brown or brown. Leg measurements: F Pt Ti Mt T I 1.55+0.12 0.94+0.05 1.30+0.12 0.95+0.07 0.62+0.05 II 1.62+0.12 0.92+0.08 1.37+0.16 0.99+0.07 0.60+0.04 III 1.49+0.12 0.79+0.05 1.02+0.08 1.17+0.10 0.53+0.07 IV 2.09+0.22 0.95+0.07 1.70+0.11 2.10+0.13 0.66+0.05 Leg armature: Ti II v.2.1 (one case), IV d.2.2, v.1.1.0 (one case), Mt III d.2.1.2 (2.0.2), lat.2.2(1.2), v.2.2. Abdomen brown, pale brown, or greyish-brown, 3.53+0.41 long, 2.10+0.29 wide, ratio 1.69+0.13. Epigyne as in Figs. 2022. DERIVATIO NOMINIS. This species is devoted to my colleague and friend, Ukrainian arachnologist Nikolai (Mykola in Ukrainian) M. Kovblyuk (Taurida National University, the Crimea). DISTRIBUTION. Ukraine: The Crimea.

LITERATURE DATA. RUSSIA. ALT [Marusik et al., 1996*]. Tuva [Mikhailov, 1992b; Logunov et al., 1998-ld; Marusik et al., 2000]. KAZAKHSTAN. VKO: Ust-Kamenogorsk [Mikhailov, 1992b], Saur Mt. Ridge [Mikhailov, 1992b; Eskov, Marusik, 1995: C. cf. dvoraki]. DISTRIBUTION. Russia S-Siberia (Altais, Tuva). EKazakhstan.

Clubiona (Clubiona) riparia L. Koch, 1866
Clubiona riparia Song et al., 1999: figs. 251GH (), 253H I (). MATERIAL. RUSSIA: Chita Area, Dahurian State Reserve (SZM): 1 , SW bank of Barun-Torei Lake, steppe, 15.06.1995, R. Dudko, 1 (SZM), N bank of Zun-Torei Lake, Kuku-Khadan Mt. and its environs, 24.06.1995; I. Lyubechanskiy & V. Smirnova; 1 (ZMMU), Kamchatka, Paratunka ca. 30 km S of Yelizovo, Betula forest, litter, under bark & on vegetation, 20.09.1992, SG; 2 (ZMMU), Maritime Prov., Lazovskiy Distr., wet meadow in Kievka River floodplain, 1.09.1981, TO; S-Kuriles: 1 (ZMMU), Iturup, Burevestnik, Betula scrub with bamboo undergrowth, litter&on vegetation, 23.098.10.1992, SG; 6 (ZMMU), Shikotan Isl., Krabozavoskoe Village, 43њ49.61N, 146њ45.44E, 1018.09.1997, leg. et det. YM. More specimens are available from the Upper Kolyma, the environs of Magadan, Kamchatka, Sakhalin, Kunashir & Paramushir isls.

Clubiona (Clubiona) pseudosaxatilis Michailov, 1992
Clubiona pseudosaxatilis Mikhailov, 1992b: 1112, figs. 34 (); C. cf. dvoraki Eskov, Marusik, 1995. MATERIAL. RUSSIA: 1 (PSU), Altais, 67 km of KoshAgach, 30.07.1939, D.E. Kharitonov. KAZAKHSTAN: 4 (PSU), 2 (ZMMU), [Dzhungarskiy Alatau Mt. Ridge], Topolevka River, near Aksu-Bas Mt., under stones, 07.1936, A.G. Ovsyannikov. More specimens from VKO were also collected.

LITERATURE DATA. RUSSIA. BShK: Shulgan-Tash Reserve [Mikhailov, 1992b; Efimik, Gulyashchikh, 1995]. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]. YaKT: Central Yakutia [Koponen, Marusik, 1992*; Marusik et al., 1993a]; Aldan River: Krestkhaldzhey [Mikhailov, 1992b]. MGD: Upper Kolyma [Marusik et al., 1992; Marusik, 1994; Mikhailov, Marusik, 1995]; environs of Magadan [Marusik et al., 1992; Mikhailov, Marusik, 1995; Bukhkalo, 2002]; Chaun Bay [Marusik et al., 1992]; Cisokhotia [Marusik et al., 1992]; Chukotka [Mikhailov, Marusik, 1995]; Anadyr [Mikhailov, 1992b]. KMCh [Mikhailov, 1992b; Mikhailov, Marusik, 1995]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Sokhondo Reserve and environs [Mikhailov, 1992b]. AMR [Mikhailov, 1992b]. KhAB: N part [Marusik et al., 1992; Mikhailov, Marusik, 1995]; S part [Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b]; Kuriles [Kamura, 2001]; S-Kuriles: Kunashir, Iturup isls. [Marusik


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UKRAINE: 7 , 7 (ZMMU), Kiev Area, ca. 40 km SSE of Kiev, right side of Kanevskoe Reservoir, 2 km S of Stayki Village, sandy bank, 1719.06.1998, YM. AZERBAIJAN: 1 (ZMMU), Kuba, park, 20.06.1976, PD; 1 (ZMMU), Nabran ca. 30 km NW of Khachmas, Quercus, Carpinus, Acer etc. forest, litter & under bark, 2122.04.1987, SG & KE.

et al., 1993b-ld]; N-Kuriles: Paramushir, Shumshu isls. [Mikhailov, 1992b; Marusik et al., 1993b-ld]. DISTRIBUTION. Russia S-Urals, C- & E-Siberia, Far East (continental & island). Mongolia, Japan (Hokkaido), China (Guizhou, Jilin, Heilongjiang), N-America.

Clubiona (Clubiona) terrestris Westring, 1851
LITERATURE DATA. ESTONIA [Vilbaste, 1993]. BELARUS: Belovezhskaya Pushcha Reserve [Mikhailov, 1992b]. UKRAINE: Volynskaya & Lvov areas [Esyunin et al., 1993, 1995; Penev et al., 1994]. MISIDENTIFICATIONS. RUSSIA. KRN: Boguchany Distr. [Izmailova, 1989b]. This record is clearly outside the known range of this species. DISTRIBUTION. Estonia, Belarus, Ukraine, Moldavia. Not found in Russia. All Europe.

REMARKS. 1. A re-examination of all the available C. similis males from the ex-USSR territory has revealed the absence of true C. similis with only one exception (Transcarpathia). C. frisia seems to be allopatric with European C. similis. The European range of C. frisia remains unclear. 2. Two females from the Crimea (Sakskiy Distr. and Krymskiy State Nature Reserve) should also belong to C. frisia. LITERATURE DATA. RUSSIA. BRN [Mikhailov, 1992b: C. similis]. KRS [Wunderlich, Schuett, 1995]. STV [Wunderlich, Schuett, 1995]. NVS [Mikhailov, 1992b: C. similis]. DISTRIBUTION. Russia European part, the Caucasus, S-Siberia. Ukraine, Azerbaijan. C-Europe, Great Britain [Merrett, Murphy, 2000].

T

HE SIMILIS - GROUP

Clubiona (Clubiona) similis L. Koch, 1867
MATERIAL. UKRAINE: 1 (NHMB), Zakarpatskaya Area, Nagy Bocskу (Mбrmaros).

Clubiona (Clubiona) neglecta O. Pickard-Cambridge, 1862 Figs. 2627.
Clubiona similis sensu Azheganova, 1951: Esyunin, Pakhorukov, 1992; C. yadongensis Hu, 2001: 291292, fig. 8-169: 12 (), syn.n.; C. xiningensis Hu, 2001: 293294, fig. 8-271: 14 (), syn.n. MATERIAL [measurements of chelicerae are given in square brackets, for details, see below]. RUSSIA: 1 , 1 (ZMMU), Ryazan Town, Oktyabrskiy Distr., slope near creek, ruderal vegetation, 0110.06.1996, leg. et det. D. Osipov; 1 [0.42] (ZMMU),North Ossetia, Sunzhenskiy Mt. Ridge, 30.0724.08.1985, S.K. Alekseev; Tuva: 1 [0.40], 1 (ZMMU), environs of Kyzyl Town, agrobiological station, 25.05.1989, DL; 1 [0.48], 4 (ZMMU), same locality, 650700 m, 01.07.1990, DL; 2 [0.41, 0.44], 22 (ZMMU), Erzin Distr., 2025 km NW of Erzin, DusKhol Lake, 800900 m, 31.05.1989, DL; 1 [0.38] (ZMMU), Erzin Distr., 30 km W of Erzin, 11001500 m, 27.05.1989, DL; 3 [0.41, 0.42, 0.45], 1 (ZMMU), environs of Kyzyl Town, right bank of Enisey River, floodplain forest, 07.06.1989, DL; 2 [0.40, 0.40], 5 (ZMMU), Tandinskiy Distr., near Chagytai Lake, steppe, Artemisia, 1100 m, 28.06.1989, DL; 1 [0.40], 1 (ZMMU), Krasnoyarsk Province, Khakassia, Altaiskiy Distr., ca. 40 km SE of Beliy Yar Town, ca. 35 km E of Novorossiyskoe, Berezovsloe Lake, 350400 m, 2223.06.1990, DL; 1 , 1 (SZM), Chita Area, Dahurian State Reserve, SW bank of BarunTorei Lake, steppe, 15.06.1995, R. Dudko. More specimens were also collected in Novosibirsk Area and Altais (SZM). GEORGIA: 1 [0.43] (ZMMU), Batumi, 10.08.1981, DL; 1 [0.32], 1 (ZMMU), Batumi, Feriya Mt., ca. 100 m, 07.11.1981, DL. AZEBAIJAN: 1 [0.38] (ZMMU), Shemakha Distr., Pirkuli Reserve, wet clearing in forest belt, 02.06.1984, DL; 1 [0.57] (ZMMU), Lerik Distr., Divagach, 2.07.1985, PD. ARMENIA: 8 (ZMMU, SZM), Sevan Town, near railway station, under stones, 28.07.1983, DL & V.I. Ovstharenko. KAZAKHSTAN: 1 (SZM), Pavlodar, 03.06.1994, OL; 1 , 5 (SZM), Taldy-Kurgan Area, Gvardeiskiy Distr., 7 km E of Kospan, 1820.06.1996, I.B. Skvorkin; 1 (ZMMU), Alma-Ata Area, 56 km SE of Bakanas, right bank of Ili River, 44њ46N, 76њ20E, 0203.06.2000, A.V. Gromov. KIRGHIZIA: 2 [0.40, 0.45] (ZMMU), Kuturga, 27.07.1977, S.L. Zonstein; 2 [0.39, 0.46], 5 (ZMMU), Kungei-Alatoo Mt. Range, Chon-Uryukty, 20002500 m, Picea forest belt, 21.06.1983, S.V. Ovchinnikov. More specimens were also available from Kirghizia. TADJIKISTAN: 1 (ZMMU), Khatlon Area, Kuybyshev Distr., right bank of Vakhsh River, floodplain, tugai community, 10.1988, A.P. Kononenko.

REMARKS. Following the recent description of the closely related species C. frisia Wunderlich et Schьtt, 1995 which differs only in the male, all the literature data based on males requires a re-examination of the pertinent specimens. All female data may belong to either C. similis or C. frisia. LITERATURE DATA (all doubtful!). RUSSIA. VRN [Mikhailov, 1992b: ; Panteleeva, Karavaev, 1992]. ChVSh [Oliger M., 1996, 1999]. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993-ld]. SMR [Krasnobaev, Matveev, 1993]: Zhiguli Reserve [Krasnobaev, 1996]. AST [Mikhailov, 1992b: ]. PRM [Esyunin, 1995; Esyunin, Efimik, 1996]. ChLB: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. Transbaikalia [Danilov, 1999-ld]. LITHUANIA [Pupiska, 1939; Vilkas, 1992-ld; Relys, 1994-ld]. UKRAINE. DNTs [Rudnikova, 1994]: Luganskiy Reserve [Prokopenko, 1999a]. Polesye [Evtushenko, 1991b, 1993b]. GEORGIA [Mkheidze, 1997]. AZERBAIJAN: Lenkoran [Guseinov, 1998, 1999]. DISTRIBUTION. Ukraine (Transcarpathia). Doubtful data: Russia: European part, Urals, Transbaikalia; Baltic countries, Ukraine, Georgia, Azerbaijan. All Europe. Mongolia (?). According to recent data, the distribution of this species in Europe is rather sporadic; it is absent from such different regions as the Netherlands [Helsdingen, 1999] and Slovakia [Gajdos et al., 1999].

Clubiona (Clubiona) frisia Wunderlich et Schьtt, 1995
Clubiona similis (part.) Mikhailov, 1990a, 1992b; C. frisia Wunderlich et Schuett, 1995: 1016, Abb. 110 (). MATERIAL. RUSSIA: 1 (ZMMU), Bryansk Area, Bezhetsk Distr., Gorodishchi, N.A. Kalinichev; 1 , 1 (ZMMU), Kazan, 06.05.1987, G.A. Pushch; 1 (ZMMU), Ulyanovsk Area, Vyazovka, 29.0405.05.1992, V. Zolotukhin; 1 (ZMMU, Stavropol Province, E of Novopavlovsk, forest of Quercus, Salix, Ulmus, etc., along stream, 28.05.1982, SG; 1 (ZMMU), Novosibirsk, Umgeb. v. Akademgorodok, 2331.05.1986, W. Schawaller.


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24 26 23

27
Figs. 2327. Clubiona spp., left palp: 2325 C. pseudoneglecta (Hungary), 2627 C. neglecta, Tuva (26), Batumi, Georgia, the Caucasus (27). 23 ventrally, 24 retrolaterally, 2527 ventro-retrolaterally. Рис. 2327. Clubiona spp., левый пальпус: 2325 C. pseudoneglecta (Венгрия), 2627 C. neglecta, Тува (26), Батуми, Грузия (27). 23 вентрально, 24 ретролатерально, 2527 вентро-ретролатерально. ROMANIA: 1 (NHMB), Hargita, 04.07.1942.

REMARKS. 1. The index of the basal cheliceral segments length / carapaces length is given in square brackets for males (see Remarks under the next species). 2. Judging from the original drawings of C. yadongensis Hu, 2001 () and C. xiningensis Hu, 2001 (), both species are undoubtedly synonymous with C. neglecta. LITERATURE DATA. RUSSIA. ChVSh [Sidorenko, Dunin, 1997; Sidorenko, 1998]. SMR [Krasnobaev, Matveev, 1993; Dunin, 2001]: Zhiguli Reserve [Krasnobaev, 1996]. PRM [Esyunin, Efimik, 1995; Pakhorukov et al., 1995]. BShK [Esyunin, Efimik, 1996; Efimik, 1997]: Bashkirian Reserve [Efimik, 1995a]; Shulgan-Tash Reserve [Efimik, Gulyashchikh, 1995]. ChLB: Ilmenskiy Rserve [Polyanin, Lagunov, 1992]; Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. ORN [Esyunin, Efimik, 1996; Efimik et al., 1997]. NVS [Mikhailov, 1992b]. ALT [Marusik et al., 1996]. KRN: Khakassia [Mikhailov, 1992b]. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]. IRK: Baikal Lake: Birkchin, Zamogoi isls. [Izmailova, 1989a]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Sokhondo Reserve and environs

[Mikhailov, 1992b]. KhAB: S part [Kim, Kurenshchikov, 1995]. ESTONIA [Vilbaste, 1974, 1993]. LITHUANIA [Pupiska, 1939; Vilkas, 1992-ld; Relys, 1994-ld; Relys, Dapkus, 2001]. UKRAINE. Polesye [Evtushenko, 1991b,c]; KhRK [Prisnyi, 1993]. GEORGIA [Mkheidze, 1997]. AZERBAIJAN [Dunin, Mamedov, 1992]: Lenkoran [Guseinov, 1998, 1999]. KAZAKHSTAN. Kora [Mikhailov, 1992b]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996-ld]. DOUBTFUL DATA. UKRAINE: DNTs: Khomutovskaya Step Reserve [Polchaninova, 1993]; Luganskiy Reserve [Prokopenko, 1999a]; steppes of Levoberezhnaya Ukraine [Polchaninova, 1996]; Askania-Nova Reserve [Polchaninova, 1995]. Should belong to C. pseudoneglecta. Material requires examination. MISIDENTIFICATIONS. UKRAINE: KhRS: Chernomorskiy Reserve [Polchaninova, 1998]; the Crimea [Mikhailov, 1992b]. All identifications actually belong to C. pseudoneglecta (material examined).


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and Tuva to Transbaikalia and Mongolia. In Central Europe, C. pseudoneglecta is found only in Germany and Switzerland, whereas C. neglecta has a considerably broader range. LITERATURE DATA. RUSSIA. KRS: environs of Krasnodar [Seifulina, 2002*]. UKRAINE: KhRS: Chernomorskiy Reserve [Polchaninova, 1998: C. neglecta]; The Crimea [Mikhailov, 1992b: C. neglecta]. DISTRIBUTION. Russia S-European part. Ukraine, Moldavia, Azerbaijan, Armenia. C-Europe, Great Britain [Merrett, Murphy, 2000], Bulgaria.

DISTRIBUTION. Russia C-European part, the Caucasus, Urals, S-Siberia. Baltic countries, Belarus, Ukraine, Moldavia, Georgia, Azebaijan, Armenia, Kazakhstan, Kirghizia, Tadjikistan. All Europe, Morocco, Afghanistan, Mongolia, China (Shansi), S-Korea.

Clubiona (Clubiona) pseudoneglecta Wunderlich, 1994 Figs. 2325.
Clubiona neglecta (part) Mikhailov, 1992b; C. pseudoneglecta Wunderlich, 1994: 157160, Abb. 13 (). MATERIAL. RUSSIA: Krasnodar Province: 1 (ZMMU), Slavyansk Distr., wheat, sweeping, 11.06.1970, N. Egorova; 2 , 1 (ZMMU), env. of Krasnodar, 10 km NW of Starokorsunskaya Settlement, 2427.06.2000, R. Seifulina. UKRAINE: 2 (ZMMU), environs (N) of Dnepropetrovsk, shore of Lenina Lake, 1819.06.1982, A.A. Zyuzin; 1 [0.75], 1 (ZMMU), Voroshilovgrad (=Lugansk) Area, Provalie, 12.06.1988, N.Yu. Polchaninova; 1 [0.69], 1 (ZMMU), Kherson Area, Golopristanskiy Distr., Rybalchie, Chernomorskiy Reserve, steppe, 1988, N.Yu. Polchaninova; 1 [0.71] (ZMMU), The Crimea, Simferopol Distr., Krasnolesie, 11.07.1981, V.A. Bragina. MOLDAVIA: 2 (ZMMU), Kagul Distr., environs of Roshu, 0506.07.1990, S.V. Makarenkov. AZERBAIJAN: 1 (ZMMU), Kutkashen Distr., Vandam, 10.08.1979, PD; 2 [0.73, 0.74] (ZMMU), Lerik Distr., Gosmalyan, 0104.07.1985, PD. ARMENIA: 2 (ZMMU, SZM), Sevan Town, near railway station, under stones, 28.07.1983, DL & V.I. Ovstharenko. HUNGARY: , (NHMB, ZMMU), different localities (for details, see Mikhailov & Szinetбr, 1997). BULGARIA: 1 (IZW), Stara Planina, Konstantin [Konstantinov] ad Tirnovo, 22.07.1964, J. Plisko.

Clubiona (Clubiona) pseudosimilis Michailov, 1990
MATERIAL. RUSSIA: 1 Suzan-Su Canyon, 06-19.07.1999, AZERBAIJAN: 2 (ZMMU), Dogguzul, 2000 m, 05.08.2001, E. (SZM), Kabardino-Balkariya, A.V. Barkalov. Gusary [former Kusary] Distr., Guseinov.

LITERATURE DATA. RUSSIA. KRS: Caucasian Reserve [Mikhailov, 1992b]. More material was collected in the same reserve (ZIN). AZERBAIJAN: Lenkoran [Guseinov, 1998, 1999]. DISTRIBUTION. Russia Caucasus Major. Georgia, Azerbaijan, Armenia.

The germanica-subgroup Clubiona (Clubiona) germanica Thorell, 1871
MATERIAL. RUSSIA: 1 (PC), Belgorod Area, 25 km W of Belgorod, environs of Kustovoe, bank of Vorskla River, Salix and dense multiherbaceous grassland, 07.10.1996 leg. et det. AP; 1 (ZMMU), Tomsk Area, 25 km of Kargasok Village, deciduous forest, under bark, 12.08.1983, AZ; 1 (ZMMU), Tuva, Biy-Khem River Basin, Serlik-Khem River, 8 km upstream of the mouth, 11.06.1992, A.B. Ryvkin. KAZAKHSTAN: Pavlodar Area: 1 (ZMMU), Uspenskiy Distr., 15 km E of Bogatyr Sovkhoz, Betula-Populus tremula groves, 19.07.1990, OL; 1 (SZM), Bayan-Aulskiy Distr., environs of Kyzyl-Tau Village, meadow, 11.06.1991, OL.

REMARKS. Differences between the females of C. neglecta and C. pseudoneglecta are obvious [s. Wunderlich, 1994]. Nevertheless, no distinguishing characters are found in the male copulatory organs. The degree of tegular sclerotization is variable and corrspondingly the seminal duct may or may not be clearly visible (cp. Figs. 2327). J. Wunderlich [pers. comm., 1996] suggested a useful character: the basal cheliceral segments length / carapaces length index. Most of the male specimens of C. neglecta and C. pseudoneglecta given above have had this index calculated (in square brackets). My data show the following indices: Tuva + Khakassia 0.380.45 (true neglecta), Kirghizia 0.390.46 (true neglecta), the Caucasus 0.380.47, 0.57 (true neglecta) and 0.73 (pseudoneglecta), Ukraine 0.690.75 (pseudoneglecta). An index of 0.57 (Azerbaijan, Divagach) marks a rather intermediate position between the two species. The record of both species in same locality of Armenia is of special zoogeographical interest. C. pseudoneglecta inhabits the steppes and forest-steppes in East Europe. C. neglecta seems to prefer meadows and also dry steppe biotopes. Both species occur sympatrically in the Caucasus (Azerbaijan, Armenia). C. neglecta seems to be absent from most of Ukraine and Moldavia, as well as from Hungary. The easternmost records of C. pseudoneglecta are East Ukraine (Provalie) and Krasnodar Province of Russia. C. pseudoneglecta and C. neglecta were found also in the collection of Prof. S.A. Spassky (ZIN), but unfortunately specimens and labels were mixed in a single vial and therefore they could not be matched properly. The records from Rostov Area (Russia) and Berdyansk (Ukraine) are tentatively assigned to C. pseudoneglecta. C. neglecta is distributed from Europe (more northern localities in East Europe) through Kazakhstan

LITERATURE DATA. RUSSIA. KRL [Wo?ny, 1995]: Kivach Reserve [Tsellarius, 1993]. PSK [Mikhailov, 1992b]. TVR [Esyunin et al., 1993, 1995; Penev et al., 1994]. MSK [Seifulina, Chernyshev, 2001]. KLZh [Mikhailov, 1992b]. VRN [Panteleeva, Karavaev, 1992]. ChVSh [Sidorenko, Dunin, 1997]. SMR [Krasnobaev, Matveev, 1993]: Zhiguli Reserve [Krasnobaev, 1996]. ARKh [Mikhailov, 1992b]: Pinezhskiy Reserve [Krasnobaev, 1990]. PRM [Pakhorukov et al., 1995; Esyunin, Efimik, 1996]. EKB [Esyunin, Efimik, 1996]: Visimskiy Reserve [Esyunin, Ukhova, 2001]. BShK: Bashkirian Reserve [Efimik, 1995a]; Shulgan-Tash Reserve [Efimik, Gulyashchikh, 1995]. ChLB [Esyunin, Efimik, 1996]: Satka Distr. [Mikhailov, 1992b]; Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. TMN: Tarko-Sale [Mikhailov, 1992b], Yuganskiy Reserve [Esyunin, 1996]. NVS [Mikhailov, 1992b]. ALT [Marusik et al., 1996*]. KRN: Tsentralnosibirskiy Reserve [Mikhailov, 1992b], environs of Mirnoe [Marusik et al., 2002b; Rybalov et al., 2002]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]. ESTONIA [Vilbaste, 1974]. LITHUANIA [Pupiska, 1939; Rakauskas, 1995; Vilkas, 1992-ld; Relys, 1994-ld]. UKRAINE. Polesye [Evtushenko, 1991b, 1993a]; ChRK [Esyunin et al., 1993, 1995; Penev et al., 1994]; DNTs: Khomutovskaya Steppe Reserve [Polchaninova, 1993].


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28

29

30

31

32

Figs. 2832. Clubiona congentilis: 2830 left palp, ventrally (28), ventro-retrolaterally (29), retrolaterally (30), 3132 genitalia, epigyne (31) and vulva (32). Рис. 2832. Clubiona congentilis: 2830 левый пальпус, вентрально (28), вентро-ретролатерально (29), ретролатерально (30), 3132 гениталии , эпигина (31) и вульва (32).

GEORGIA [Mkheidze, 1997]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996-ld]. DISTRIBUTION. Russia C- & N-European part, W& C-Siberia. Baltic countries, Belarus, Ukraine, Georgia, Kazakhstan, Kirghizia. S-, C- & N-Europe.

Clubiona (Clubiona) congentilis Kulczyсski, 1913 Figs. 2832.
MATERIAL. RUSSIA: 1 (PSU), Orenburg Area, Sol-Iletsk Distr., Shybyndy Ravine, forest, 22.01.2001, S.L. Esyunin.

UKRAINE: 1 , 4 (PC), 2 , Kharkov Area, Zmiev Distr., Gaidary Village, 1973, A.V. Prisnyi. KAZAKHSTAN: 1 (SZM), Akmola Area, near Atbasar Town, 0117.06.1997, U.V. Gromova, N.A. Chernova, A.V. Gromov; Pavlodar Area: 1 (ZMMU), Bayan-Aulskiy Distr., Babaly Mts., stony debris, 27.08.1990, OL; 1 (SZM), same distr., environs of Kyzyl-Tau Settlement, meadow, 11.06.1991, OL; 1 (ZMMU), 25 km N of Pavlodar, floodplain of Irtysh River, Populus-Salix forest, clearings, 06.06.1992, OL. For additional specimens see Mikhailov [1992b].

DIAGNOSIS. This species is closest to C. germanica, but differs in the following characters: the embolus occupies the upper half of the bulb (first third in C. germanica) and is


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arched around it (acutely entangled in C. germanica); the long and thin copulatory tubes and the copulatory openings are separated and situated close to the epigynal median line (fused in a single groove in C. germanica). DESCRIPTION. MALE (8). Carapace 2.46+0.47 long, 1.74+0.32, ratio 1.42+0.26. Carapace, chelicerae and legs reddish-coloured or pale. Leg measurements: F Pt Ti Mt T I (8) 1.77+0.260.93+0.14 1.58+0.28 1.23+0.26 0.79+0.15 (11) 1.77+0.461.02+0.22 1.49+0.29 1.10+0.29 0.76+0.16 II (8) 1.82+0.310.94+0.14 1.62+0.27 1.25+0.19 0.81+0.13 (11) 1.80+0.421.01+0.24 1.52+0.29 1.14+0.21 0.77+0.17 III (8)1.60+0.240.77+0.101.17+0.261.36+0.23 0.56+0.06 (11) 1.60+0.350.86+0.22 1.12+0.22 1.30+0.36 0.58+0.10 IV (8)2.07+0.340.91+0.141.69+0.24 2.15+0.36 0.72+0.11 (11) 2.14+0.580.99+0.25 1.71+0.33 2.17+0.38 0.72+0.17 Leg armature: F III d.1.1.3 (1.2.3), Pt III rlat.1 (0), Ti III d.2.2, v.1.1.1 (1.1), Mt III d.2.1.2 (2.2), lat.1.2 (2.2), v.2.2. Palp as in Figs. 2830. Lengths of F, Pt. Ti, and Cb are 0.73+0.12 (7), 0.40+0.07 (7), 0.30+0.03 (6), 0.92+0.15 (6), respectively. Abdomen brown or dark brown, 2.74+0.70 long, 1.79+ 0.19 wide, ratio 1.65+0.17. FEMALE (11). Carapace 2.72+0.60 long, 1.96+0.52 wide, ratio 1.40+0.20. Carapace reddish-coloured or brown; chelicerae chestnut-coloured, rarely dark brown, brown or reddish-coloured; legs pale, reddish-coloured, rarely brown. Leg measurements see above. Leg armature as in male, but Pt III rlat.1, Ti II v.2.2 (2, 1.2), III d.2.2, v.1.1. Abdomen dark brown or brown, 4.23+1.71 long, 2.54+ 1.09 wide, ratio 1.68+0.20. Epigyne as in Figs. 3132. LITERATURE DATA. RUSSIA. MSK [Mikhailov, 1992b]. BLG [Mikhailov, 1992b]. MRS [Krasnobaev, Matveev, 1993]. RST [Mikhailov, 1992b]. ORN [Esyunin et al., 1999*]. KAZAKHSTAN. SKO [Mikhailov, 1992b]. PVL [Mikhailov, 1992b]. AKM [Mikhailov, 1992b]. VKO [Mikhailov, 1992b]. DISTRIBUTION. Russia, Ukraine, Kazakhstan. Steppes and forest-steppes from the former Czechoslovakia (exact locality unknown: Mikhailov, 1992b) to E-Kazakhstan.

T

HE OBESA - GROUP

Clubiona (Clubiona) furcata Emerton, 1919
LITERATURE DATA. RUSSIA. YaKT: Khandyga [Mikhailov, 1992b; Marusik et al., 1993a-ld]. MGD: Upper Kolyma [Marusik et al., 1992-ld; Marusik, 1994; Mikhailov, Marusik, 1995]; Chukotka [Marusik et al., 1992]. Transbaikalia [Danilov, 1999-ld]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Siberia, N-Far East (continental). N-America.

Clubiona (Clubiona) kularensis Marusik et Koponen in Marusik et al., 2002
Clubiona kularensis Marusik et Koponen in Marusik et al., 2002a: 357, figs 17 (). MATERIAL. RUSSIA: 1 (ZMMU), Khabarovsk Province, Pravaya Bureya River upper flow, mountain tundra, 30.06.2000, L.A. Trilikauskas.

DISTRIBUTION. Russia Priamurie (new data), NE Yakutia.

Clubiona (Clubiona) kurilensis Bцsenberg et Strand, 1906
Clubiona kurilensis Hayashi, Chikuni, 1984: 15, figs. 110 (); Hayashi, 1987: figs. 12 (); Paik, 1990b: 6971, figs. 1927 (); Song et al., 1999: 425, figs. 246WX (), 250AB (). MATERIAL. RUSSIA: Kamchatka: , (ZMMU, SZM), Dolina Geizerov, different biotopes, 30.085.09.1987, AT; 1 (ZMMU), Petrovavlovsk-Kamchatskiy, Zavoikovo Distr., 17.09.1996, leg. et det. YM; 5 , 4 (OC, ZMMU), Maritime Prov., Lazovskiy Distr., 19761984, TO; 4 , 10 (ZMMU, SZM), N-Kuriles, Paramushir Isl., SW shore, Shelikhova Bay, 50њ22N, 155њ37E, 1325.08.1996, leg. et det. YM; 1 (ZMMU), S-Kuriles, Khabomai Archipelago, Zelyonyi Isl., W coast of Srednee Lake, Drepanocladus-Aulacomium bog with Carex, Lysiachiton and dwarfish Salix, 6.08.1994, KE. More specimens were collected in Kunashir & Iturup isls.

T

HE JAPONICOLA - GROUP

Clubiona (Clubiona) japonicola Bцsenberg et Strand, 1906
Clubiona japonicola Gong, 1984: 203208, figs. 1630 (); Mikhailov, 1990b: 151152, figs. 2324 (); Paik, 1990b: 6567, figs. 112 (); Song et al., 1999: 416, figs. 246LM (), 249KL (). MATERIAL. RUSSIA: 1 inad, 2 juv. (ZMMU), Maritime Prov., Dalnevostochnyi Marine Reserve, Stenina Isl., lake, 30.06.1997, leg. T. Vshivkova, det. YM. More specimens were collected in PRI: Khanka Lake & env. of Vladivostok Town. N-KOREA: 2 , 2 (ZIN), N Pyongan Prov., env. of Sinanchzhu, 09.07.1950, leg. N. Borkhsenius, det. D.E. Kharitonov.

LITERATURE DATA. RUSSIA. AMR [Mikhailov, 1992b]. KhAB: S part [Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b], N- & S-Kuriles [Kamura, 2001]; Kunashir & Iturup Isls. [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia S-Far East (continental & island) + N-Kuriles & Kamchatka. Japan, Korea, China (widely [Song et al., 1999]).

Clubiona (Clubiona) logunovi Michailov, 1990
Clubiona logunovi Mikhailov, 1990b: 154155, figs. 2526 ().

LITERATURE DATA. RUSSIA. SAKh: S-Kuriles [Kamura, 2001] material not examined. DISTRIBUTION. Russia S-Far East (continental & ? island). Japan, N- & S-Korea, China (widely [Song et al., 1999]), Taiwan, Philippines, India [Majumder & Tikader, 1991, as C. bengalensis Biswas, 1984].

LITERATURE DATA. RUSSIA. KhAB: Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995-ld]. PRI: SikhoteAlin Reserve [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (continental).

Clubiona (Clubiona) marusiki Michailov, 1990
DISTRIBUTION. Known only from Russia: AMR (type locality).
Clubiona marusiki Mikhailov, 1990b: 155, figs. 2728 ().


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34

Figs. 3334. Clubiona latericia, genitalia (Maritime Prov., Khanka Lake shore), under KOH-clearing: 33 epigyne, 34 vulva. Рис. 3334. Clubiona latericia, гениталии (Приморье, берег оз. Ханка), просветлено в KOH: 33 эпигина, 34 вульва.

Clubiona (Clubiona) paiki Michailov, 1991
Clubiona paiki Mikhailov, 1991: 227, figs. 5456 (); C. chechtsirica Mikhailov, 1995a: 102, figs. 46 (), syn.n. MATERIAL. RUSSIA: 1 , 1 (ZMMU), Maritime Prov., Sikhote-Alinskiy Reserve, Kordon Kabany, 45њ0816N, 135њ52 40E, 650900 m, taiga, 30.0604.07.1999, leg. Yu. Sundukov, det. YM (as C.ch.).

REMARKS. The synonymy of C. chechtsirica under C. paiki is obvious from the the single sample of both the male and female from the same habitat. LITERATURE DATA. RUSSIA. KhAB: S part [Kim, Kurenshchikov, 1995: C.p.; Kurenshchikov, 1997a,b: C.ch. + C.p.]: Bolshekhekhtsyrskiy Reserve [Mikhailov, 1995a: C. ch.]. DISTRIBUTION. Russia S-Far East (continental).

Clubiona (Clubiona) praematura Emerton, 1909
LITERATURE DATA. RUSSIA. YaKT: Lower Kolyma [Marusik et al., 1993a-ld; Mikhailov, Marusik, 1995]. MGD: Chukotka [Marusik et al., 1992; Marusik, 1993; Mikhailov, Marusik, 1995]. DISTRIBUTION. Russia N-Far East (continental). N-America.
MATERIAL. More specimens were collected in Chukotka.

The latericia-subgroup Clubiona (Clubiona) latericia Kulczyсski, 1926 Figs. 3334.
Clubiona latericia Mikhailov, 1990b: 155156, figs. 2932 (). MATERIAL. RUSSIA: 1 (ZMMU), Krasnoyarsk Province, W part of Taimyr Peninsula, ca. 9 km W of Pyasina Lake NW coast, 0110.07.2000, D. Osipov;Maritime Prov.: 1 (ZMMU), Lazovskiy Reserve, Petrova locality, wet meadow, on soil, 03.08.1980, TO; 1 , 2 (ZMMU), Khanka Lake, CE shore, 44њ39N, 132њ34E, 1516.07.1998, leg. et det. YM; 3 , 4 (ZMMU), N-Kuriles, Paramushir Isl., SW shore, Taina River, 50њ22N, 155њ36.67E, 21.08.1996, leg. et det. YM; 1 (ZMMU), S-Kuriles, Khabomai

REMARKS. After KOH-clearing, the copulatory openings become visible (Figs. 3334). In earlier descriptions [Mikhailov, 1990], they were not indicated due to obscurity (darkness) of this part of the epigyne. Unfortunately, the shape of the spermathecae appears to be incorrect (disrupted) following exposure to KOH. LITERATURE DATA. RUSSIA. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]. YaKT: Central Yakutia [Koponen, Marusik, 1992*; Marusik et al., 1993a-ld]; Dulgalakh River [Marusik et al., 1993a]; Lower Kolyma [Marusik et al., 1992; Marusik et al., 1993a; Mikhailov et al., 1995]; Moyero River [Mikhailov, Marusik, 1995]; Lower Yana River [Marusik et al., 2002a]. MGD: Upper Kolyma [Marusik et al., 1992-ld; Marusik et al., 1994; Mikhailov, Marusik, 1995]; Chukotka [Marusik et al., 1992ld; Mikhailov, Marusik, 1995]; Chaun Bay [Marusik et al., 1992]. KMCh [Mikhailov, Marusik, 1995]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. KhAB: Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995ld]. SAKh: Sakhalin [Marusik et al., 1993b]; N- & S-Kuriles [Kamura, 2001]. DISTRIBUTION. Russia S- & E-Siberia, Far East (continental & island). Alaska.

Archipelago, Zelyonyi Isl., W coast of Srednee Lake, Drepanocladus-Aulacomium bog with Carex, Lysiachiton and dwarfish Salix, 06.08.1994, KE. More specimens were collected in Tuva and Bolshekhekhtsirskiy Reserve.

Clubiona (Clubiona) kunashirensis Michailov, 1990
Clubiona kunashirensis Mikhailov, 1990b: 156157, figs. 33 37 (); C. lyubarskii Mikhailov, 1991: 227228, figs. 5759 (); C. kunashirensis Ono et al., 1991: figs. 3031 (); Mikhailov, 1992b: 15 (= C. lyubarskii), 1995a: 103 (= C. sharkeyi); C. sharkeyi Hayashi, 1994: 5859, figs 79 (). MATERIAL. RUSSIA: 1 (ZMMU), Sakhalin, SW part, Krilyon Peninsula, E side, Uryum River ca. 3 km from its mouth, 46њ28.078N, 142њ19.746E, 1718.08.2001, YM.


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38

35

36

37

Figs. 3538. Clubiona yaroslavi sp.n., holotype, left palp: 35 subventrally, 36 ventro-prolaterally, 37 ventro-retrolaterally, 38 retrolaterally. Рис. 3538. Clubiona yaroslavi sp.n., голотип, левый пальпус: 35 субвентрально, 36 вентро-пролатерально, 37 вентроретролатерально, 38 ретролатерально. More specimens were collected in Iturup and Kunashir isls.

LITERATURE DATA. RUSSIA. SAKh: S-Kuriles [Kamura, 2001]; Kunashir Isl. [Marusik et al., 1993b-ld], Iturup Isl. [Mikhailov, 1992b; Marusik et al., 1993b-ld], Moneron Isl. [Marusik et al., 1993b]. DISTRIBUTION. Russia Sakhalin (new data), SKuriles. Japan: Hokkaido.

Clubiona (Clubiona) yaroslavi Michailov, sp.n. Figs. 3538.
MATERIAL. Holotype (ZMMU Ta-6596), Maritime Prov., Siniy Gai, forest, litter, 14.06.1991, A.A. Borok.

Clubiona (Clubiona) subborealis Michailov, 1992
Clubiona subborealis Mikhailov, 1992b: 1516, figs. 56 (). MATERIAL. RUSSIA: 1 (ZMMU), Buryatia, Barguzin Mt. Ridge, Olso River, 54њ52N, 110њ55E, 950 m, 06.07.1996, SK.

LITERATURE DATA. RUSSIA. Transbaikalia [Danilov, 1999]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. KhAB: S part [Kurenshchikov, 1999 *]; Komsomolsk Reserve [Mikhailov, 1992b; Kim, Kurenshchikov, 1995-ld]. DISTRIBUTION. Russia the mountains of South Siberia from Transbaikalia to Priamurie.

Clubiona (Clubiona) tsurusakii Hayashi, 1987
Clubiona tsurusakii Hayashi, 1987: 3940, figs. 2330 (). MATERIAL. RUSSIA: S-Kuriles: 1 (ZMMU), Kunashir Isl., CW part, 44њ00.38N, 145њ40.95E, sulphur springs, 05.09.1997, YM.

LITERATURE DATA. RUSSIA. SAKh: Kunashir Isl. [Mikhailov, 1992b; Marusik et al., 1993b-ld]. DISTRIBUTION. Russia S-Kuriles: Kunashir Isl. Japan.

DIAGNOSIS. This species belongs to the obesa-group (C. logunovi, C. tateyamensis Hayashi, 1989, and C. tsurusakii) which is characterized by a protruding embolar region of peculiar shape (cp. Hayashi, 1987: fig. 28; 1989: fig. 18; Mikhailov, 1990b: fig. 25) and by the moderately attenuating embolus starting at the middle, not at the apical part of EP with an apical 1/51/3 rectangled and laying on the membraneous tegular conductor (cp. also Mikhailov, 1995b). This species differs from other congeners by having a unifurcate TA (widened in other species). DESCRIPTION. MALE. Body length 4.75. Carapace 2.18 long, 1.48 wide, ratio 1.47. Carapace and chelicerae reddish straw-coloured. Palp as in Figs. 3538. Lengths of F, Pt, Ti, and Cb are 0.70, 0.30, 0.28, and 0.85, respectively. Legs straw-coloured. Leg measurements: F Pt Ti Mt T I 1.83 0.75 1.83 1.38 0.75 II 1.88 0.83 1.80 1.38 0.78 III 1.65 0.75 1.18 1.43 0.55 IV 2.30 0.85 1.88 2.25 0.98 Leg armature: F III d.1.1.3. Abdomen straw-coloured, 2.65 long, 1.30 wide, ratio 2.04. FEMALE unknown. DERIVATIO NOMINIS. The species name is devoted to my son Yaroslav (born 1995), a good field naturalist and spider-collector.


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Figs. 3940. Clubiona microsapporensis, genitalia (Ussuri Reserve): 39 epigyne, 40 vulva. Рис. 3940. Clubiona microsapporensis, гениталии (Уссурийский заповедник): 39 эпигина, 40 вульва.

40

The sapporensis-subgroup Clubiona (Clubiona) sapporensis Hayashi, 1986
Clubiona sapporensis Hayashi, 1986: 3537, figs. 1017 (); C. sapporensis Mikhailov, 1990b: 157158, figs. 3844 (). MATERIAL. RUSSIA: 8 , 5 (OC, ZMMU), Maritime Prov., Lazovskiy Distr. and Reserve, 19751984, TO; 1 , 2 (ZMMU), Sakhalin Area, Moneron Isl., SSE shore, 24.06.2001, leg. et det. YM; S-Kuriles: 5 (SZM), 27 (ZMMU, SZM), Khabomai Archipelago, Zelyonyi Isl., W coast 1.5 km N of Glushnevskiy Cape, shingle-sandy seashore, under wracks and driftwoods, 06.08.1994, KE; 1 , 3 (ZMMU), Shikotan Isl., Delfin Bay, seashore, under stones and driftwoods, 11.08.1994, KE; 2 (ZMMU), same island, E coast, small bay opposite to Isl. of Greag, 43њ48.50N, 146њ47.19E, 16.07.1997, YM. More specimens were collected in PRI: Khasan Distr., Kunashir & Iturup isls.

REMARKS. Some unusual specimens of C. charitonovi cannot be distinguished from C. sapporensis by the wide interval between the copulatory openings; however the 2nd spermathecae are always longer in the former species than in the latter. DISTRIBUTION. Russia S-Siberia, S-Far East (continental & island).

Clubiona (Clubiona) eskovi Michailov, 1995
LITERATURE DATA. RUSSIA. PRI: Lazovskiy Reserve [Mikhailov, 1995a]. DISTRIBUTION. This species is known only from the type locality.
Clubiona eskovi Mikhailov, 1995a: 103105, figs. 1516.

LITERATURE DATA. RUSSIA. AMR: Khingan Reserve [Mikhailov, 1992b]. SAKh: C- & S-Kuriles [Kamura, 2001]; Kunashir Isl. [Marusik et al., 1993b-ld]; Iturup Isl. [Mikhailov, 1992b; Marusik et al., 1993b-ld]; Urup Isl. [Marusik et al., 1993b]. DISTRIBUTION. Russia S-Far East (continental & island). Japan: Hokkaido; N-Korea. ECOLOGY. In coastal regions, this species occurs in sandy and intertidal shore biotopes.

Clubiona (Clubiona) microsapporensis Michailov, 1990 Figs. 3940.
Clubiona microsapporensis Mikhailov, 1990b: 158159, figs. 4849 (). MATERIAL. RUSSIA: Maritime Prov.: 4 (OC), Lazovskiy Distr., 19761983, TO; 1 (ZMMU), Ussuri Reserve, 43њ39N, 132њ33E, 2931.07.1998, YM; 6 (ZMMU), Khasan Distr., environs of Andreevka Village, 42њ3536N, 131њ13E, 1115.08. 1998, YM; 1 (ZMMU), Sakhalin, central part, Leonidovka River (right tributary of Poronai River), 49њ15.092N, 142њ43.889E, 0405.08.2001, YM.

Clubiona (Clubiona) charitonovi Michailov, 1990
Clubiona charitonovi Mikhailov, 1990b: 159160, figs. 4547 (), 5051 (). MATERIAL. RUSSIA: 1 (SZM), SE Tuva, ca. 610 km N of Shuurmak, Larix-Betula forest, sweeping on glades, 1000 m, 10.07.1993, DL. More specimens are also available from KhAB: Bolshekhekhtsirskiy Reserve.

LITERATURE DATA. RUSSIA. KRN: Minusinsk [Mikhailov, 1992b]. IRK [Mikhailov, 1992b]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Daurskiy Reserve [Mikhailov, 1992b]. AMR [Mikhailov, 1992b]. KhAB: Sredne-Tambovskoe [Mikhailov, 1992b]; Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995-ld]. SAKh: S-Kuriles [Kamura, 2001]; Kunashir Isl. [Mikhailov, 1992b; Marusik et al., 1993b-ld].

REMARKS. 1. The epigyne structure is variable: sometimes the vulva is longer than wide (the specimen from NKorea), or the 2nd spermathecae are not tubular, but almost rounded, as in Figs. 3940 (the specimen from Ussuri Reserve). 2. In some cases, the body length exceeds the limits indicated in the original description. Another distinguishing character is proposed, viz., the copulatory openings in C. microsapporensis are directed anteriorly, whereas those in C. sapporensis though directed anteriorly, are also inclined to the median line of the epigyne (cp. figs. 48 and 42 in Mikhailov, 1990b). In addition, the epigynal area of C. microsapporensis is paler in colour than in C. sapporensis. LITERATURE DATA. RUSSIA. KhAB: Amur River [Mikhailov, 1992b]; Slavyanka [Mikhailov, 1992b; Kim, Kurenshchikov, 1995]; Bolshoi Khekhtsir Mt. Ridge [Kim,


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Figs. 4142. Clubiona dunini sp.n., paratype, genitalia: 41 epigyne, 42 vulva. Рис. 4142. Clubiona dunini sp.n., паратип, гениталии : 41 эпигина, 42 вульва.

42

Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: SKuriles [Kamura, 2001]. DISTRIBUTION. Russia S-Far East (continental). N-Korea.

1992b]. PRI [Mikhailov, 1992b]: Kedrovaya Pad Reserve [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (continental). S-Korea, China (Liaonin, Jilin).

Clubiona (Clubiona) nenilini Michailov, 1995
LITERATURE DATA. RUSSIA. S-Siberia [Mikhailov, 1995a]. DISTRIBUTION. Exact locality remains unknown.
Clubiona nenilini Mikhailov, 1995a: 102103, figs. 1314 ().

Clubiona (Clubiona) kurenshikovi Michailov, 1995
LITERATURE DATA. RUSSIA. KhAB: S part [Kurenshchikov, 1997b]; Bolshekhekhtsirskiy Reserve [Mikhailov, 1995a]; Bolshoi Khekhtsir Mt. Ridge [Kurenshchikov, 1997a]. DISTRIBUTION. This species is known only from the type locality.
Clubiona kurenshikovi Mikhailov, 1995a: 102, figs. 1112 ().

The corrugata-subgroup Clubiona (Clubiona) corrugata Bцsenberg et Strand, 1906
Clubiona corrugata Hu, Song, 1983: 55, fig. 1AD (); Gong, 1984: 202203, figs. 115 (); Song et al., 1999: 415, figs. 245DE (), 247MN (). MATERIAL. RUSSIA: 1 , 2 (ZMMU), Maritime Prov., Khanka Lake CE shore, 44њ39N, 132њ34E, 1516.07.1998, leg. et det. YM; S-Kuriles, Shikotan Isl.: 1 (ZMMU), Ostrovnaya River 500 m upsteram off mouth, wet meadow of Carex & Lysiachiton with Salix bushes, 11.08.1994, KE; 1 (ZMMU), Krabozavodskoe Village, 43њ49.61N, 146њ45.44E, 1018.09.1997, YM.

The bakurovi-subgroup Clubiona (Clubiona) bakurovi Michailov, 1990
Clubiona bakurovi Mikhailov, 1990b: 163164, figs. 6165 (); C. bakurovi Song et al., 1999: 415, figs. 247EF (). MATERIAL. RUSSIA: 1 , 2 (OC), Maritime Prov., Lazovskiy Distr., Tachinghou locality, mixed forest, N slope, 10.09. 1980, TO. Additional specimens were collected in PRI: Anisimovka (Kangauz) and Chernyshevka.

LITERATURE DATA. RUSSIA: PRI [Marusik, Koponen, 2000]. DISTRIBUTION. Russia S-Far East (continental & island). Japan, Korea, China (widely [Song et al., 1999]).

LITERATURE DATA. RUSSIA. KhAB: Bolshoi Khekhtsir Mt. Ridge [Mikhailov, 1992b; Kim, Kurenshchikov, 1995-ld]; Slavyanka [Kim, Kurenshchikov, 1995-ld]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b]. DISTRIBUTION. Russia S-Far East (continental & island). N-Korea. China: Jilin.

Clubiona (Clubiona) kimyongkii Paik, 1990
Clubiona kimyongkii Paik, 1990a: 9899, figs. 5965 (); C. ussurica Mikhailov, 1990b: 161162, figs. 5253 (), 5860 (); C. kimyongkii Song et al., 1991: 6869, fig. 3AD (, = ussurica); 1999: 425, figs. 246ST (), 249NO (). MATERIAL. RUSSIA, Maritime Prov.: 3 , 2 (CO, ZMMU), Lazovskiy Reserve, 19751984, TO; 2 (SZM), 20 km SE of Ussuriysk Town, Gornotaezhnaya Village, mixed mountainous forest, 0102.06.1994, V.V. Dubatolov.

Clubiona (Clubiona) dunini Michailov, sp.n. Figs. 4142.
MATERIAL. Holotype (ZMMU Ta-6597), Russia, S part Maritime Prov., 43њ34N, 134њ12E, Oblachnaya Mt., 1400 03.08.1998, YM. Paratypes: Maritime Prov.: 5 (ZMMU, JWC), same data holotype; 2 (ZMMU, JWC), 43њ06N, 132њ48E, environs Anisimovka Village, Litovka Mt., 1200 m, 26.07.1998, YM. of m, as of

LITERATURE DATA. RUSSIA. AMR: Khinganskiy Reserve [Mikhailov, 1992b]. KhAB: Slavyanka [Mikhailov, 1992b; Kim, Kurenshchikov, 1995]; Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995]; Bikin Distr. [Mikhailov,

DIAGNOSIS. This species is most closely related to C. bakurovi, but differs in having well-divided copulatory openings, the epigyne less sclerotized posteriorly, and the position of the 2nd part of the spermatheca differs. The special depressions for fixation of the male tibial apophysis are also absent.


The spider genus Clubiona in the fauna of the former USSR: 2003 update
DESCRIPTION. FEMALE (8). Body length 6.25+1.69. Carapace 2.74+0.04 long, 2.01+0.03 wide, ratio 1.36+0.02. Carapace and legs straw-coloured. Chelicerae straw- or reddish-straw-coloured, 1.01+0.02 long. Leg measurements: F Pt Ti Mt T I 1.85+0.06 1.01+0.02 1.53+0.05 1.13+0.03 0.75+0.01 II 1.96+0.07 1.03+0.03 1.63+0.06 1.18+0.06 0.74+0.01 III 1.71+0.08 0.87+0.01 1.21+0.03 1.42+0.08 0.64+0.01 IV 2.40+0.10 1.03+0.02 1.90+0.08 2.44+0.18 0.76+0.04 Leg armature: Ti III d.2.2, v.1.1.0 (1.1.1), IV d.2.2, v.1.1.1 (2.1.1, 1.2.1), Mt IV d.2.1.2, lat.2.2, v.2.1.2 (2.0.2). Abdomen (pale) brown, 3.62+1.29 long, 2.11+1.28 wide, ratio 1.75+0.19. Epigyne as in Figs. 4142. MALE unknown. DERIVATIO NOMINIS. This species is devoted to my late colleague and friend, the arachnologist Dr. Peter M. Dunin (19521998) who made important contributions to the study of spiders of the Russian Far East.

303

The irinae-subgroup Clubiona (Clubiona) irinae Michailov, 1991
Clubiona irinae Mikhailov, 1991: 208210, figs. 17 (); C. nigra Zhang, 1993: 162163, figs. 14 (); C. irinae Mikhailov, 1995a: 105 (= C. nigra); Song et al., 1999: 416, figs. 246HI (), 249 GH (). MATERIAL. RUSSIA: 1 (ZMMU), Khabarovsk Province, Ussuri River, Khoperskoe Village, 10.07.1931, V.I. Sytshevskaya; 6 , 12 (OC, ZMMU), Maritime Prov., Lazovskiy Distr. and Reserve, 19751979, 1981, 1983, 1984, TO. More specimens are also collected in PRI: environs of Vladivostok and Ussuri Reserve.

LITERATURE DATA. RUSSIA. KhAB: S part [Kim, Kurenshchikov, 1995]; Komsomolsk Reserve [Mikhailov, 1992b]; Bolshoi Khekhtsir Mt. Ridge [Kurenshchikov, 1999]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b]. DISTRIBUTION. Russia S-Far East (continental & island). N-Korea, NE-China (Jilin, Heilongjiang).

[Matveev, 1991; Krasnobaev, Matveev, 1993]. ChVSh [Sidorenko, Dunin, 1997]. ULN [Mikhailov, 1992b]. TTR: Volzhsko-Kamskiy Reserve [Aleynikova et al., 1979]. SMR [Mikhailov, 1992b; Krasnobaev, Matveev, 1993; Dunin, 2001]. RST [Minoranskiy, 1995]. PRM [Pakhorukov et al., 1995; Esyunin, Efimik, 1996]. EKT [Esyunin, Efimik, 1996]. BShK [Esyunin & Efimik, 1996; Efimik, 1997]: Bashkirian Reserve [Ghirfanova et al., 1992; Efimik, 1995a]. ChLB [Esyunin, Efimik, 1996]: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. ORN [Esyunin, Efimik, 1996; Efimik et al., 1997]. KRG [Mikhailov, 1992b]. NVS [Mikhailov, 1992b]. KRN: environs of Mirnoe [Marusik et al., 2002b; Rybalov et al., 2002]; W Sayan Mts. [Logunov et al., 1998*]. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]. ALT [Marusik et al., 1996]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]: Dzherghinskiy Reserve [Danilov, 1995]. ChIT [Mikhailov, 1992b]. KhAB: S part [Kim, Kurenshchikov, 1995]. SAKh: Sakhalin [Mikhailov, 1992b; Marusik et al., 1993b-ld]. ESTONIA [Vilbaste, 1974: as C holosericea L.]. LITHUANIA [Pupiska, 1939: asC holosericea L.; Rakauskas, 1985; Vilkas, 1992-ld; Relys, 1994-ld; Relys, Dapkus, 2001]. UKRAINE. ODS [Esyunin et al., 1993, 1995; Penev et al., 1994]. Polesye [Evtushenko, 1991a,b,c, 1993a,b]. KhRK [Polchaninova, Prisada, 1995]. DNTs [Rudnikova, 1994, 1995]: Luganskiy Reserve [Prokopenko, 1999a]. KhRS: Chernomorskiy Reserve [Polchaninova, 1998], Askania-Nova Reserve [Polchaninova, 1995]. ZhTM: Polesskiy Reserve [Guryanova, 1994]. GEORGIA [Mkheidze, 1997]. KAZAKHSTAN. PVL [Mikhailov, 1992b]. DOUBTFUL RECORD. RUSSIA. YaKT [Belimov, 1975]. DISTRIBUTION. Russia European part, Caucasus, W- & S-Siberia, S-Far East (except Maritime Prov.). Baltic countries, Belarus, Ukraine, Moldavia, Georgia, Azerbaijan, Armenia, Kazakhstan. All Europe, Mongolia, N-America.

Clubiona (Clubiona) ezoensis Hayashi, 1987
Clubiona ezoensis Hayashi, 1987: 3638, figs. 1522 (); 1994: figs 36 (); C. ezoensis Mikhailov, 1991: 211212, figs. 816 (). MATERIAL. RUSSIA: S-Kuriles: , (ZMMU, SZM), Iturup Isl., 1992, SG, 1994, KE. One more specimen was collected in Kunashir Isl.

T

HE PALLIDULA - GROUP

Clubiona (Clubiona) pallidula (Clerck, 1758)
MATERIAL. RUSSIA: 2 (PC), Belgorod Area, environs of Borisovka and Shebekino, 1986, 1997, AP; 1 (ZMMU), Chelyabinsk Area, Yuzhnouralsk, Kichighinskiy Bor forest,Betula stand with young Pinus, 04.07.1991, KM; 1 (ZMMU), 80 km E of Kemerovo, Zolotoi Kitat River valley, 31.05.1995, YM. KAZAKHSTAN, Pavlodar Area (all leg. by OL): 1 (ZMMU), Uspenskiy Distr., 15 km E of Bogatyr Sovkhoz, Betula-Populus tremula groves, 19.07.1990; 1 (SZM), Lebyazhinskiy Distr., 2 km NW of Chernoe Village, Irtysh River floodplain, 21.09.1990; 2 (ZMMU, SZM), same distr., near Malybay Lake, 16.05.1991; 1 (ZMMU), Bayal-Aulskiy Distr., environs of Kyzyl-Tau Village, meadow, 11.06.1991. More specimens were collected in Samara and Pavlodar areas (Russia & Kazakhstan).

LITERATURE DATA. RUSSIA. KhAB: Kizi Lake [Mikhailov, 1992b]. SAKh: Sakhalin [Mikhailov, 1992b; Marusik et al., 1993b-ld], N- & S-Kuriles [Kamura, 2001]; Iturup Isl. [Marusik et al., 1993b], Kunashir Isl. [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia S-Far East (continental & island). Japan: Hokkaido.

Clubiona (Clubiona) haeinsensis Paik, 1990
Clubiona haeinsensis Paik, 1990a: 9193, figs. 1631 (); C. haeinsensis Song et al., 1999: 416, figs. 245T, 246A ().

LITERATURE DATA. RUSSIA. LNN: Nizhnesvirskiy Reserve [Oliger T., 1996]. MSK [Seifulina, Chernyshev, 2001]. KLZh [Esyunin et al., 1993]. TLS [Esyunin et al., 1993]. RZN: Okskiy Reserve [Esyunin et al., 1993]. BLG: Les-Na-Vorskle Reserve [Esyunin et al., 1993, 1995; Penev et al., 1994]. VRN [Panteleeva, Karavaev, 1992]. MRS

LITERATURE DATA. RUSSIA. AMR: Khinganskiy Reserve [Mikhailov, 1995a]. KhAB: Bolshekhekhtsirskiy Reserve [Mikhailov, 1995a]. DISTRIBUTION. Russia S-Far East (continental). S-Korea, China (Hubei).


304

K.G. Mikhailov Clubiona (Clubiona) komissarovi Michailov, 1992
Clubiona komissarovi Mikhailov, 1992b: 1920, fig. 7 (). MATERIAL. RUSSIA: 2 (OC), Maritime Prov., Lazovskiy Reserve, Sukhoi Klyuch, valley forest 21.06.1981, TO.

LITERATURE DATA. RUSSIA. PRI [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East: Maritime Prov. N-Korea.

Clubiona (Clubiona) langei Michailov, 1991
MATERIAL. RUSSIA: 4 (OC, ZMMU), Maritime Prov., Lazovskiy Distr., 19771983, TO. One more specimen was collected in PRI: Anisimovka Village. (). Clubiona langei Mikhailov, 1991: 213214, figs. 2223

44

LITERATURE DATA. RUSSIA. KhAB: Troitskoe [Mikhailov, 1992b], Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995-ld]. PRI [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (continental).

Clubiona (Clubiona) mandschurica Schenkel, 1953

43
Figs. 4344. Clubiona helenae sp.n., holotype, left palp: 43 ventro-retrolaterally, 44 tibial apophysis. Рис. 4344. Clubiona helenae sp.n., голотип, левый пальпус: 43 вентро-ретролатерально, 44 вырост голени.

Clubiona mandschurica Zhu, Yu, 1982: 61, fig. 2ad (); Paik, 1990a: 105107, figs. 101112 (); Ono, 1994: 3940, figs. 34 (); Mikhailov, 1995a: 105106, figs. 17 19 (); 1997b: 190191, figs. 12 (); Song et al., 1999: 425, fig. 252KL (). MATERIAL. RUSSIA: 1 (ZMMU), Maritime Prov., Lazovskiy Reserve, Korpad locality, 01.06.1980, TO.

DISTRIBUTION. First record for Russia. Japan (Honsu), N- & S-Korea, China (Jilin, Shaanxi, Beijing).

Clubiona (Clubiona) helenae Michailov, sp.n. Figs. 4344.
MATERIAL. Holotype (ZMMU Ta-6598), Russia, Maritime Prov., Lazovskiy Reserve, Sukhoy Klyuch Camp, plain forest, 24.07.1980, TO.

Clubiona (Clubiona) oligerae Michailov, 1995
Clubiona oligerae Mikhailov, 1995a: 108, figs. 2832 (). MATERIAL. RUSSIA, Maritime Prov.: 1 (ZMMU), Merkushevka, forest litter, 20.07.1991, A.A. Borok; 1 (ZMMU), Kamenushka, 16.07.2001, YM.

DIAGNOSIS. The new species is very close to C. oligerae Michailov, 1995, but differs in possessing a spine-like tooth on the TA and it does not have a dark inner side of the EP (cp. fig. 28 in Mikhailov, 1995a). DESCRIPTION. MALE. Body length 5.30. Carapace 2.38 long, 1.80 wide, ratio 1.32. Carapace, chelicerae and legs reddish. Palp as in Figs. 43-44. Lengths of F, Pt, Ti, and Cb are 0.75, 0.38, 0.30, and 0.88, respectively. Leg measurements: F Pt Ti Mt T I 1.75 0.88 1.83 1.33 0.83 II 1.88 0.93 1.85 1.35 0.83 III 1.65 0.78 1.25 1.55 0.63 IV 2.15 0.93 2.00 2.43 0.75 Leg armature: F III d.1.1.3, III d.1.2.3 (1.3.3). Abdomen greyish-reddish, 3.13 long, 1.45 wide, ratio 2.16. FEMALE unknown. DERIVATIO NOMINIS. This species is dedicated to my wife, Elena (Helen in English) Mikhailova, an arachnologist.

REMARKS. The basal width of the TA is wider than that figured in the origial description (Fig. 31 in Mikhailov, 1995a). Fig. 32 in that paper is more correct. LITERATURE DATA. RUSSIA. PRI: Lazovskiy Reserve [Mikhailov, 1995a]. DISTRIBUTION. Russia Maritime Province.

Clubiona (Clubiona) phragmitis C.L. Koch, 1843
MATERIAL. RUSSIA, Maritime Prov., Khanka Lake: 1 , 4 (ZMMU), CE shore, 44њ39N, 132њ34E, 1516.07.1998, YM; 1 (ZMMU), CW shore, Sosnovyi Isl. & pnsl. nearby, 44њ52N, 132њ07E, 17.08.1998, YM. KAZAKHSTAN, Pavlodar Area: 1 (SZM), Lebyazhinskiy Distr., 2 km NW of Chernoe Village, Irtysh River floodplain, 21.09.1990, OL; 1 (ZMMU), 30 km SE of Pavlodar, steppe, waterbodies, 21.10.1990, OL. KIRGHIZIA: 3 (ZMMU, SZM), environs of Frunze (=Bishkek), Karagachovaya Roshcha, 1979, S.L. Zonshtein.

LITERATURE DATA. RUSSIA. LNN: Nizhnesvirskiy Reserve [Oliger T., 1996]. PSK [Mikhailov, 1992b]. MSK [Seifulina, Chernyshev, 2001]. MRS [Matveev, 1991; Kras-


The spider genus Clubiona in the fauna of the former USSR: 2003 update
nobaev, Matveev, 1993-ld; Matveev et al., 1995]. ChVSh [Oliger M., 1996, 1999]. SMR [Krasnobaev, Matveev, 1993]: Zhiguli Reverve [Krasnobaev, 1996]. TMB [Mikhailov, 1992b]. ARKh [Mikhailov, 1992b]. PRM [Esyunin et al., 1993; Pakhorukov et al., 1995]. EKB [Esyunin, Efimik, 1996]. ChLB: Troitskiy Reserve [Esyunin, Pakhorukov, 1992], Ilmenskiy Reserve [Esyunin, Efimik, 1996]. BShK [Efimik, 1997]: Shulgan-Tash Reserve [Efimik, Gulyashchikh, 1995], Bashkirian Reserve [Efimik, 1995a]. Tuva [Mikhailov, 1992b; Logunov et al., 1998-ld; Marusik et al., 2000]. It is obvious from the newly studied material (see above) that earlier references to AMR [Azheganova, Stenchenko, 1977] and PRI [ternbergs, 1988], which were previously suggested to be erroneous [s Mikhailov, 1992b], may actually be valid. A re-examination of the pertinent specimens is necessary. ESTONIA [Vilbaste, 1974]. LITHUANIA [Pupiska, 1939; Vilkas, 1992-ld; Relys, 1994; Relys, Dapkus, 2001]. UKRAINE. North Prichernomorye/ Cis-Chernomoria [Mikityuk, 1991]. Environs of Berdyansk [Mikhailov, 1992b]. DNTs: Khomutovskaya Step Reserve [Polchaninova, 1993]. Askania-Nova Reserve [Polchaninova, 1995]. Polesye [Evtushenko, 1991b,c, 1993a]. MOLDAVIA [Ro?ca, 1941]. AZERBAIJAN. Lenkoran [Guseinov, 1998, 1999]. KAZAKHSTAN. Cisaralia [Mikhailov, 1992b]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996-ld]. DISTRIBUTION. Russia all European part, Caucasus, mountains of Middle Asia, S-Siberia, S-Far East (rarely). Baltic countries, Belarus, Ukraine, Moldavia, Azerbaijan, Kazakhstan, Kirghizia. S-, C- & N-Europe, Iran [Mozaffarian et al., 1998, 2000*], China (Qinghai, Xinjiang).

305

According to the body size and distribution data, the female of C. langei (length of carapace 4.05+0.86) is probably conspecific with the male of C. zacharovi (length of carapace 3.503.85). A detailed ecological study is required. LITERATURE DATA. RUSSIA. PRI [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East: Maritime Prov. N-Korea [Mikhailov, 1997b].

Clubiona (Clubiona) zyuzini Michailov, 1995
LITERATURE DATA. RUSSIA. PRI: Lazovskiy Reserve [Mikhailov, 1995a]. DISTRIBUTION. This species is known only from the type locality.
Clubiona zyuzini Mikhailov, 1995a: 106108, figs. 2627 ().

The hummeli-subgroup Clubiona (Clubiona) odesanensis Paik, 1990 Figs. 4546.
Clubiona odesanensis Paik, 1990a: 9698, figs. 4758 (); C. odesanensis Mikhailov, 1991: 212213, figs. 1721 (); Song et al., 1999: 426, figs. 250OP (), 253B (). MATERIAL. RUSSIA: 1 (SZM), Amur Area, Blagoveshchensk, 08.1996, E.I. Malikova; Maritime Prov.: 13 , 1 (OC, ZMMU), Lazovskiy Distr., 19781984, TO; 1 , Lazovskiy Reserve, Korpad Camp, 43њ16N, 134њ08E, 0609.08.1998, YM. One more specimen is available also from PRI: Ussuri Reserve.

Clubiona (Clubiona) sopaikensis Paik, 1990
Clubiona sopaikensis Paik, 1990a: 9394, figs. 3239 (); C. sopaikensis Mikhailov, 1991: 214215, figs. 2429 (). MATERIAL. RUSSIA: Maritime Prov.: 1 (ZMMU), S Sikhote-Alin Mt. Range, Gorelaya Sopka Mt., 43њ3030N, 134њ06 08E, 13001470 m, 1720.06.1999, Yu. Sundukov; 2 , 1 (OC), Lazovskiy Distr., Kievka, grass, sweeping, 10.07.1982, TO. More specimens were collected in Khasan Distr.

LITERATURE DATA. RUSSIA. AMR: Khinganskiy Reserve [Mikhailov, 1992b]. KhAB: S part [Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia S-Far East (continental & island). N- & S-Korea.

Clubiona (Clubiona) zacharovi Michailov, 1991
Clubiona zacharovi Mikhailov, 1991: 226, figs. 5253 (). MATERIAL. RUSSIA, Maritime Prov.: 1 (OC), Lazovskiy Reserve, Petrova locality, forest edge, 06.06.1978, TO; 5 (OC, ZMMU), Lazovskiy Reserve, 1980, 1981, TO; 2 , Ussuri Reserve, 43њ39N, 132њ33E, 2931.07.1998, SK&YM; 1 (ZMMU), W of Khanka Lake, environs of Barabash-Levada Village, 44њ45.5N, 131њ27E, 1819.07.1998, leg. et det. YM.

REMARKS. In one of the specimens from Lazovskiy Reserve the tibial apophysis is smooth and the tooth-like process is not visible. The whole TA is not always so flat as is clear from the original description [Mikhailov, 1991: fig. 5253].

REMARKS. 1. The very similar females of C. odesanensis and C. sopaikensis can be distinguished by the atria, which in the former species are positioned more laterally relative to the copulatory openings. 2. The male of C. odesanensis is probably conspecific with the female of C. papillata. In all the material of C. odesanensis examined by me since 1990, the males of C. odesanensis and the females identified as C. papillata have been found together (i.e. in the same samples) 7 times, while only 3 samples have contained both the males and females of C. odesanensis (including type series). It is peticularly interesting that most of the records of C. odesanensis females are from Korea, whereas the males are confined mostly to the Russian Far East. For example, the type series of this species (S-Korea) includes 14 females and only 1 male. In NKorea, 5 males and 5 females were collected. In Russia, 23 males and only 2 females were caught! See also comments under C. papillata. 3. The male of C. odesanensis is very similar to C. hummeli Schenkel, 1936. Re-examination of a fragment of the C. hummeli lectotype from Naturhistorisches Museum Basel (Figs. 4749) revealed several distinguishing characters in TA structure, especially in dorsal view (cp. Figs. 46 and 47). In C. hummeli, the TA dorsal branch is bilobate at less than a half of its length, or, at least, bifurcate in certain view. In C. odesanensis, it is unambiguously solid. The ventral surface of the TA dorsal branch is flattened, concave, smooth and semi-transparent in C. odesanensis, whereas it bears an additional process in C. hummeli. The double ridge on the EP apophysis is longer in C. odesanensis, covering the base of the embolus in retrolatero-ventral view. LITERATURE DATA. RUSSIA. KhAB: S part [Mikhailov, 1992b; Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (continental). N- & S-Korea, China (Jilin).


306

K.G. Mikhailov

45 46

47

48

49

Figs. 4549. Clubiona spp., right palp, tibial apophysis: 4546 C. odesanensis (Maritime Prov., Kievka), 4749 C. hummeli (part of lectotype, Naturhistorisches Museum Basel). 45, 48 retrolaterally, 46, 47 dorsally, 49 ventro-retrolaterally. Рис. 4549. Clubiona spp., правый пальпус. вырост голени: 4546 C. odesanensis (Приморье, Киевка), 4749 C. hummeli (фрагмент лектотипа, Естественноисторический музей, Базель). 45, 48 ретролатерально, 46, 47 дорсально, 49 вентроретролатерально.

Clubiona (Clubiona) papillata Schenkel, 1936 Figs. 5053.
Clubiona papillata Schenkel, 1936: 162163, fig. 56a-b (); Clubiona wolchongsensis Paik, 1990a: 99100, figs 6673 (), syn.n.; C. wolchongsensis Mikhailov, 1991: 225226, figs. 5051 (); Song et al., 1991: 71, fig. 6AB (), 1999: 427, fig. 252CD (); C. serrulata Hu et al., 1991: 47, figs. 58 (), syn.n.; C. flexa Zhang, 1992: 5354, figs. 1316 (, ); C. flexa Zhang, Chen, 1993: 306308, figs 14 (, );

C. papillata Mikhailov, 1998: 89, figs.78 (,= flexa); Song et al., 1999: 426, fig. 250ST (); C. flexa Song et al., 1999: 415, figs. 245NO (), 248HI (); C. serrulata Songet al., 1999: 427, figs. 251KL (), 253LM (). MATERIAL. RUSSIA, Maritime Prov.: 16 (OC, ZMMU, SZM), Lazovskiy Distr. & Reserve, 19761982, 1984, TO; 2 (ZMMU), environs of Vladivostok, Okeanskaya Station, 24.07.1994, KE; 5 (ZMMU, ZMT), Ussuri Reserve, 43њ39N, 132њ33E, 29 31.07.1998, SK & YM; 1 (ZMMU), environs of Anisimovka Village, 43њ10N, 132њ46E, pitfall traps, 28.0709.08.1998, YM; 1 (ZMMU), Uglovaya Village, 43њ21N, 132њ03E, 04.10.1997, leg. et det. YM.


The spider genus Clubiona in the fauna of the former USSR: 2003 update

307

50

51

52

53

Figs. 5053. Clubiona papillata, genitalia (vulva) from Middle Bikin River (50), Uglovaya Village (51), Ussuri Reserve (52), Nizhnespasskoe Village (53). Рис. 5053. Clubiona papillata, гениталии (вульва), Средний Бикин (50), дер. Угловая (51), Уссурийский заповедник (52), дер. Нижнеспасское (53).

LITERATURE DATA. RUSSIA. KhAB: Amur River from Pashkovo to Khabarovsk [Kim, Kurenshchikov, 1995*]; Vyatskoe [Mikhailov, 1992b]. PRI [Mikhailov, 1992b]. REMARKS. 1. Figures of the vulvae of four specimens from different parts of the Russian Far East illustrate a transition in all distinguishing characters between C. papillata and C. wolchongsensis (initial direction of the copulatory tubes, the shape of dark structures, etc. see Figs. 5053). Unfortunately, the direction of the copulatory tubes in the type of C. papillata is incorrectly shown in Mikhailov [1998, fig. 8]. In wolchongsensis, the light area on the epigyne is situated above the copulatory openings, whereas in true papillata it is situated more laterally. However, I consider this character not distinctive enough for distinguishing the two species. 2. The male of C. flexa (not seen) is possibly identical to C. odesanensis or C. hummeli. A more detailed study of the C. flexa types (not accessible at present) is required. A series of the C. odesanensis males examined by me showed variation both in size of the genitalia and in shape of the TA. Examination of more Chinese material is required before a firm conclusion can be drawn.

3. Judging from the figures of C. serrulata from Henan, China made by Song et al. [1999], this species is clearly a junior synonym of C. papillata. Unfortunately, the original description was not available to me, but I understand that the drawings provided by Song et al. [1999] were simply copied from it. DISTRIBUTION. Russia S-Far East. N- & S-Korea, China: Liaonin, Jilin, Heilongjiang, Gansu, (?) Shenxi, Hebei, Hubei, Henan.

The propinqua-subgroup Clubiona (Clubiona) propinqua L. Koch, 1879
Clubiona propinqua Holm, 1973: 104, figs. 106108 (); Mikhailov, 1991: 215217, figs. 3033 (); 1992b: 20 (= C. subinteriecta); 1997b: figs. 56 (); C. subinteriecta Strand, 1907 Mikhailov, 1991: 217218, figs. 3438 (). MATERIAL. RUSSIA: 4 (SZM), Altai, ca. 60 km W of KoshAgach, Taltura (Chagan-Uzun) River canyon, 22002300 m, stony debris on the upper border of Larix forest, 2530.06.1999,


308

K.G. Mikhailov Clubiona (Clubiona) pseudogermanica Schenkel, 1936
Clubiona pseudogermanica Schenkel, 1936: 5556, fig. 53a,b (); C. hummeli Schenkel, 1936: 161-162, fig. 55c,d (, not ); C. propinqua (nec L. Koch, 1879, misidentified) Hu, Song, 1983: 56, fig. 3AC (); Paik, 1990a: 102103 (, = C. salictum, ); Zhang, 1991: 1011, figs. 58 (, = C. hummeli, ); C. salictum Namkung, Kim, 1987: 2328, figs. 111 (); C. hummeli Paik, 1990a: 107108, figs. 113120 ( ); Mikhailov, 1995a: 108109, figs. 3334 (); C. salictum Ono, 1993: figs. 910 (); C. pseudogermanica Ono, 1993 (? = C. salictum); Hayashi, 1994: 6063, figs. 1621 (, = C. salictum); Mikhailov, 1995a: 109 (), 1998: 8992, figs. 912 (); Song et al., 1999: 426427, figs. 251CD (), 253DE (). MATERIAL. RUSSIA: 3 (ZMMU), Maritime Prov., Khanka Lake CNW shore, 45њ12N, 131њ59.71E, 1718.07.1998, YM.

DL; Buryatia: 1 (ZMMU), Barguzin Mt. Ridge, Olso River, 54њ52N, 110њ55E, 950 m, 05.07.1996, SK; 1 (ZMMU), Barguzin River valley, Zugdeli, 55њ01N, 111њ18E, 650 m, 09.07.1996, SK. Khabarovsk Province: 1 (ZMMU), Verkhnebureinskiy Distr., 190 km SW of Bureinskiy Reserve, mouth of Usammakh Stream (confluent of Bureya River), pebble, under stones, 09.10.1999, leg. L.A. Trilikauskas. Maritime Prov.: 1 (ZMMU), S Sikhote-Alin Mt. Range, Gorelaya Sopka Mt., 43њ3030N, 134њ0608E, 1300 1470 m, 1720.06.1999, Yu. Sundukov; 1 , 6 (OC, ZMMU), Lazovskiy Distr. and Reserve, 19781984, TO; 1 , 5 (ZMMU), Lazovskiy Reserve, Korpad Camp, 43њ16N, 134њ08E, 0609.08.1998, leg. et det. YM; 4 , 37 (ZMMU, SZM), Pravaya Izvilinka River, 43њ5255N, 134њ1723E, 0102.08.1998, leg. et det. YM; 2 , 3 (ZMMU), Oblachnaya Mt., 43њ34N, 134њ12E, 16001750 m, 03.08.1998, leg. SK, det YM. More specimens were collected in MGD: Chukotka, Amur Area, Khabarovsk Province (env. of Chegdomyn), and PRI: environs of Anisimovka Village & Ussuri Reserve.

LITERATURE DATA. RUSSIA. KRN: Ermakovo Distr. [Mikhailov, 1992b]. YaKT: Central, East Yakutia [Marusik et al., 1993a; Mikhailov, Marusik, 1995]. MGD: environs of Magadan Town [Marusik et al., 1992-ld; Mikhailov, Marusik, 1995]; Upper Kolyma [Marusik et al., 1992-ld; Marusik, 1994; Mikhailov, Marusik, 1995; Bukhkalo, 1997]; Chukotka [Marusik et al., 1992, Marusik, 1993; Mikhailov, Marusik, 1995; Mikhailov, 1997b]. KMCh [Mikhailov, Marusik, 1995]. IRK [Mikhailov, 1992b]. Transbaikalia [Danilov, 1999-ld]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. AMR [Mikhailov, 1992b]. KhAB: Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995: C.p. + C. subinteriecta]; Bureinskiy Reserve [Trilikauskas, 2001]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia Siberia E of Yenisey, all Far East (continental & island). N-Korea.

REMARKS. Figures of C. propinqua male by Paik [1990a] are inadequate for determining with certainty to which species C. mayumiae or C. pseudogermanica it belongs. Zoogeographical data suggest the latter species. LITERATURE DATA. RUSSIA. PRI [Mikhailov, 1995a]. DISTRIBUTION. fUSSR S-Far East: Maritime Prov. Japan: (Honshu), N- & S-Korea, China (widely).

T

HE RECLUSA - GROUP

Clubiona (Clubiona) reclusa O. Pickard-Cambridge, 1863
MATERIAL. RUSSIA: 2 (PC), Belgorod Area, near Borisovka, Les-Na-Vorskle Reserve, edge of Vorskla River floodplain, meadow, 03.06, 11.11.1996, leg. et det. AP; 2 (SZM), Altai Province, near Volchikha bank of Zolotoe Lake, 10.07.1999, G.N. Azarkina, A.Yu. Chuikova; 1 (ZMMU), Krasnoyarsk Province, S part, near Tanzybei, forest research station, 53њ08N, 92њ53E, 0203.06.1995, YM; 1 , 1 (ZMMU), Yakutia, Lyampeshka, forest, meadow, 17.07.1977, SK. UKRAINE: 1 (ZMMU), the Crimea, environs of Kerch, 15.06.1990, OL. KAZAKHSTAN: 1 (ZMMU), Pavlodar Area, Mayskiy Distr., environs of Kirovskiy Sovkhoz, multiherbaceous meadow, 10.08.1989, OL.

Clubiona (Clubiona) mayumiae Ono, 1993
Clubiona propinqua (nec L. Koch, 1879, misidentified) Hayashi, 1987: 35, figs. 311 (); C. pseudogermanica (nec Schenkel, 1936, misidenified) Mikhailov, 1991: 219220, figs. 3943 (), 1992a,b; Marusik et al., 1993b; Kim, Kurenshchikov, 1995; C. mayumiae Ono, 1993: 9092, figs. 18 (); Hayashi, 1994: 5960, figs 1015 (); C. mayumiae Mikhailov, 1995a: 109; 1997b: figs. 78 (). MATERIAL. RUSSIA: Kamchatka: 1 (ZMMU), environs of Milkovo, Kamchatka River floodplain, fern-multiherbaceous Betula forest, sweeping, 24.08.1987, AT; 2 , 1 (ZMMU), Petropavlovsk-Kamchatskiy, multiherbaceous-gramineous Betula forest, 1619.09.1988, AT; 3 , 1 (ZMMU), Paratunka, gramineous Betula forest, 12.08.1988, AT; 3 , 4 (ZMMU), N-Kuriles, Paramushir Isl., SW shore, different localities, 10.08 15.09.1996, YM; , (ZMMU, SZM), S-Kuriles, Iturup Isl., different localities, 1992, 1994, SG, KE. More specimens were collected in Bolshekhekhtsirskiy Reserve and Kunashir Isl.

REMARKS. The epigyne shape is variable: all internal structures can be situated close to the epigastric furrow (rare) or at a distance from it (typical, as figured in all relevant papers). LITERATURE DATA. RUSSIA. KMCh [Mikhailov, 1997b]. KhAB: S part [Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993bld]; Kuriles [Kamura, 2001]; S-Kuriles: Iturup Isl. [Marusik et al., 1993b], Kunashir Isl. [Marusik et al., 1993b-ld]; NKuriles: Shumshu Isl. [Mikhailov, 1992b]. DISTRIBUTION. Russia S-Far East (continental & island), N-Kuriles, Kamchatka. Japan: Hokkaido.

LITERATURE DATA. RUSSIA. KRL: Kivach Reserve [Tsellarius, 1993]. LNN: Nizhnesvirskiy Reserve [Oliger T., 1995, 1996]. MSK [Seifulina, Chernyshev, 2001]: PriokskoTerrasnyi Reserve [Rybalov, 1991]. KLZh [Mikhailov, 1992b]. VRN [Panteleeva, Karavaev, 1992]. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993-ld]. ChVSh [Sidorenko, Dunin, 1997; Sidorenko, 1998]. ARKh [Mikhailov, 1992b]: Pinezhskiy Reserve [Krasnobaev, 1990]. PRM [Esyunin, Efimik, 1996]. BShK [Efimik, 1997]: Bashkirian Reserve [Efimik, 1995a]. TMN: S-Yamal [Korobeinikov, Esyunin, 1995; Esyunin, 1999]; Yuganskiy Reserve [Esyunin, 1996]. KRN: environs of Mirnoe [Marusik et al., 2002a; Rybalov et al., 2002]; Khakassia [Mikhailov, 1992b]. Tuva: Azas Reserve [Krasnobaev, 1994]. KMR [Romanenko, 1998]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]. ESTONIA [Vilbaste, 1974]. LITHUANIA [Pupiska, 1939; Mikhailov, 1992b; Vilkas, 1992-ld; Relys, 1994b-ld]. UKRAINE. DNTs: Luganskiy Reserve [Prokopenko, 1999a]. Polesye [Evtushenko, 1991c, 1993b]. DISTRIBUTION. Russia European part, W- & SSiberia. Baltic countries, Lithuania, Belarus, Ukraine, N- & E-Kazakhstan. S-, C- & N-Europe.


The spider genus Clubiona in the fauna of the former USSR: 2003 update Clubiona (Clubiona) bashkirica Michailov, 1992
Clubiona bashkirica Mikhailov, 1992b: 2627, figs. 1213 (). C. sp. Esyunin et al., 1993. MATERIAL. 1 (SZM), Novosibirsk Area, Baraba Steppe, Kuibyshevskiy Distr., Zonovo Vill. (7 km N of Sergovka Vill.), middle flow of Om River, mesophytous meadow, 20.06.1961, V. Mordkovich & I. Stebaev.

309

LITERATURE DATA. RUSSIA. Urals [Esyunin, Efimik, 1997]. BShK [Efimik, 1995b, 1997]: Bashkirian Reserve [Mikhailov, 1992b; Efimik, 1995a]; Shulgan-Tash Reserve [Mikhailov, 1992b; Efimik, Gulyashchikh, 1995]. DISTRIBUTION. Russia S-Urals, West Siberia (new record).

Clubiona (Clubiona) evoronensis Michailov, 1995
LITERATURE DATA. RUSSIA. KhAB: Evoron Lake [Mikhailov, 1995a]. DISTRIBUTION. This species is known only from the type locality.
Clubiona evoronensis Mikhailov, 1995a: 109, figs. 3536 ().

Clubiona (Clubiona) interjecta L. Koch, 1879
Clubiona interjecta Holm, 1973: 104, figs. 102105 (); Song et al., 1999: 416, figs. 246FG (), 249EF (). MATERIAL. RUSSIA: 1 (ZMMU), Tomsk Area, 25 km of Kargasok Village, deciduous forest, under bark, 12.08.1983, AZ; 1 (ZMMU), environs of Magadan, Klyopka Village, 08.1994, leg. N.N. Shumatbaeva, det. YM; 4 , 3 (ZMMU), Maritime Prov., Khanka Lake CE, CNW and CW shores, 1518.07.1998, leg. et det. YM. MONGOLIA: 1 (ZMMU), Selenga Aimak, Yцrdц, 24.07.1999, M. Uhruh.

LITERATURE DATA. RUSSIA. TMN: Novyi Urengoi, Tarko-Sale [Mikhailov, 1992b]. ALT [Marusik et al., 1996]. KRN: S part [Mikhailov, 1992b]. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]. YaKT: Zhigansk, Sangar, Aldan River: Krestkhaldzhey [Mikhailov, 1992b]; Central Yakutia [Koponen, Marusik, 1992*; Mikhailov, Marusik, 1995]. MGD: Upper Kolyma [Marusik et al., 1992; Marusik, 1994; Mikhailov, Marusik, 1995; Bukhkalo, 1997, 2002]; environs of Magadan [Marusik et al., 1992]; Cisokhotia [Marusik et al., 1992]. KMCh [Marusiket al., 1992; Mikhailov, Marusik, 1995]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]: Dzherghinskiy Reserve [Danilov, 1995]. ChIT [Mikhailov, 1992b]: Dahurian Reserve [Mikhailov, 1992b]. AMR [Mikhailov, 1992b]. KhAB: S part [Mikhailov, 1992b; Kim, Kurenshchikov, 1995]. PRI [Mikhailov, 1992b]. SAKh: Sakhalin [Marusik et al., 1993b-ld]. DISTRIBUTION. Russia all Siberia, Far East (continental & island). Mongolia, China (Jilin, Heilongjiang, Sichuan).

LITERATURE DATA. RUSSIA. ARKh: Pechora River Delta [Mazura, 2000]. KOM [Mikhailov, 1992b]. PRM [Mikhailov, 1992b; Pakhorukovet al., 1995; Esyunin, Efimik, 1996; Esyunin, 1999]. EKT [Esyunin, Efimik, 1996]: Visimskiy Reserve [Ukhova, Esyunin, 1996; Esyunin et al., 2000; Ukhova, 2001]. BShK [Efimik, 1997]: Bashkirian Reserve [Efimik, 1995]; Shulgan-Tash Reserve [Efimik, Gulyashchikh, 1995]. ChLB [Esyunin, Efimik, 1996]: Ilmenskiy Reserve [Polyanin, Lagunov, 1992]. TMN: S-Yamal [Mikhailov, 1992b; Korobeinikov, Esyunin, 1995; Esyunin, 1999]; Yuganskiy Reserve [Esyunin, 1996]. NVS [Mikhailov, 1992b]. TMS [Romanenko, Lukyantsev, 1995]. KMR [Romanenko, 1998]. ALT [Mikhailov, 1992b; Marusik et al., 1996]. KRN: Stolby Reserve [Mikhailov, 1992b]; TsentralnoSibirskiy Reserve and environs [Mikhailov, 1992b]; localities along Yenisei River (5768.5њN) [Marusik et al., 2002b; Rybalov et al., 2002]; Ermakovo Distr. [Mikhailov, 1992b]; Khakassia [Mikhailov, 1992b], West Sayan Mts. [Logunov et al., 1998*]. Tuva [Mikhailov, 1992b; Logunov et al., 1998*; Marusik et al., 2000]: Azas Reserve [Krasnobaev, 1994]. YaKT [Koponen, Marusik, 1992*; Marusik et al., 1993a; Mikhailov, Marusik, 1995]: Zhigansk [Mikhailov, 1992b]. MGD: Upper Kolyma [Marusik et al., 1992-ld; Bukhkalo, 1994, 1995, 1997; Marusik, 1994; Mikhailov, Marusik, 1995]; environs of Magadan [Marusik et al., 1992; Mikhailov, Marusik, 1995]; Cisokhotia [Marusik et al., 1992-ld]. KMCh [Mikhailov, Marusik, 1995]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. KhAB: N part [Marusik et al., 1992-ld; Mikhailov, Marusik, 1995]; Badzhalskiy Mt. Ridge [Mikhailov, 1992b; Kim, Kurenshchikov, 1995]. SAKh: Sakhalin [Mikhailov, 1992b; Marusik et al., 1993b]. DISTRIBUTION. Russia N-European part, Urals, all Siberia, N- (and mountainous) Far East. N- & E-Kazakhstan. Boreomontane. Central Europe (sporadic), North Europe. Japan: Hokkaido; N-Korea; N-America from Alaska to Colorado and N-Carolina, USA.

KAZAKHSTAN: 2 (ZMMU), Pavlodar Area, environs of Zhelezenka Village, Irtysh River floodplain, Populus-Salix forest with blackberry, 06.08.1988, OL. More specimens are also available from Tuva.

Clubiona (Clubiona) maracandica Kroneberg, 1875
LITERATURE DATA. UZBEKISTAN: Samarkand [Mikhailov, 1992b]. DISTRIBUTION. This species is known only from the type locality.
Clubiona maracandica Mikhailov, 1992b: 2324, fig. 8 ().

Clubiona (Clubiona) norvegica Strand, 1900
LITERATURE DATA. RUSSIA. LNN: Nizhnesvirskiy Reserve [Oliger T., 1996]. MSK [Mikhailov, 1992b]. ARKh [Mikhailov, 1992b]. PRM [Esyunin, Efimik, 1995]. TMN: Yamal [Mikhailov, 1992b; Esyunin, 1999]; Polar Urals [Koponen et al., 1998]; Yuganskiy Reserve [Mikhailov, 1992b; Esyunin, 1996]. LITHUANIA: Zhuvintas Reserve [Mikhailov, 1992b]. DISTRIBUTION. Russia N-European part, N parts of Urals & W-Siberia. Lithuania. Boreo-montaneous, sphagnicolous, i.e., confined to the Sphagnum bogs. C- & N-Europe, N-America from Alaska to Utah and New York, USA.

Clubiona (Clubiona) kulczynskii Lessert, 1905
MATERIAL. RUSSIA: 2 (ZMMU), Kemerovo Area, Kuznetskiy Alatau Reserve, delta of Malaya Bezymyanka Stream, ca. 19 km SSW of Belogorsk, Abies-Pinus taiga forest, 500 m, 27 31.08.1993, SG & V.G. Gratshev; 3 (SZM), Altai Province, Teletskoe Lake, Artybash, 27.05.1967, A.P. Kononenko; Krasnoyarsk Province: 1 (ZMMU), Taimyr Region, down Yenisei River, Potapovo Vill., 68.50њN, 86.25њE, 08.1995, leg. L.B. Rybalov, det. YM; 2 (SZM), Bolshemurtinskiy Distr., Yukseevo Village, right bank of Yenisey River, 0607.1991, A.V. Gurov; 2 (ZMMU), Kamchatka, Dolina Geizerov, alder forest with ferns, slope, litter, 31.08.1987, AT; 1 (ZMMU), Buryatia, Barguzin Mt. Ridge, Olso River, 54њ52N, 110њ55?E, 950 m, 06.07.1996, leg. SK, det. YM.


310

K.G. Mikhailov

56 54 55

Figs. 5456. Clubiona sichotanica sp.n., holotype, left palp: 54 ventrally, 55 retrolaterally, 56 tibial apophysis, retrolaterally. Рис. 5456. Clubiona sichotanica sp.n., голотип, левый пальпус: 54 вентрально, 55 ретролатерально, 56 вырост голени, ретролатерально.

Clubiona (Clubiona) rybini Michailov, 1992
Clubiona rybini Mikhailov, 1992b: 2425, figs. 910 (); C. maracandica Zonshtein et al., 1996. MATERIAL. KIRGHIZIA: 1 (ZMMU), At-Bashinskiy Distr., At-Bashi River floodplain near At-Bashi Village, tugai vegetation, 16.06.1987, Ark.A. Shileiko & M.E. Chernyakhovskiy; 2 (ZMMU, SZM), Kochkorskiy Distr., left river bank opposite of Kochkorka Village, under stones, 24.06.1987, Ark.A. Shileiko & M.E. Chernyakhovskiy; 1 (ZMMU), Kungey-Alatoo Mt. Ridge, environs of Kiz-Kus Village, Boomskoe Canyon, 28.06.1991, S.V. Ovchinnikov. One additional specimen was collected from Kirghizia.

Clubiona (Clubiona) sichotanica Michailov, sp.n. Figs. 5456.
MATERIAL. Holotype (ZMMU Ta-6599), Russia, Maritime Prov., Sikhote-Alinskiy Reserve, Zabolochennaya River, confluence with Serebryannaya River, 21.05.1994, T. Vshivkova.

LITERATURE DATA. KAZAKHSTAN: environs of Alma-Ata [Mikhailov, 1992b]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996ld: C. maracandica + C. rybini]. DISTRIBUTION. Mountains of Middle Asia: Kazakhstan, Kirghizia.

Clubiona (Clubiona) saurica Michailov, 1992
LITERATURE DATA. KAZAKHSTAN: Saur Mt. Ridge [Mikhailov, 1992b; Eskov, Marusik, 1995]. DISTRIBUTION. This species is known only from the type locality.
Clubiona saurica Mikhailov, 1992b: 2526, fig. 11 (); C. cf. maracandica Eskov, Marusik, 1995.

DIAGNOSIS. Differs from the members of the reclusagroup by the following characters: the relatively simple, not branched, embolus (resembling that in C. kulczynskii); the absence of embolic processes; the relatively short protector, not exceeding the limits of the alveolum; dorsal branch of TA with angled tip. DESCRIPTION. MALE. Body length 5.60. Carapace 2.60 long, 1.80 wide, ratio 1.44, reddish. Chelicerae reddishbrown, 1.10 long. Palp as in Figs. 5456. Lengths of F, Pt, Ti, and Cb are 0.80, 0.35, 0.28, and 0.75, respectively. Legs straw-coloured. Leg measurements: F Pt Ti Mt T I 2.45 1.20 2.25 1.80 0.93 II 2.28 1.20 2.25 1.80 0.93 III 1.88 0.90 1.40 1.70 0.58 IV 2.45 1.00 2.00 2.63 0.75 Leg armature Ti IV d.2.2, v.1.1.1 (2.1.1). Abdomen pale brown with dark brown lanceolate pattern, 3.20 long, 1.63 wide, ratio 1.97.


The spider genus Clubiona in the fauna of the former USSR: 2003 update
FEMALE unknown. DERIVATIO NOMINIS. This species is named after the Sikhote-Alin Mt. Ridge (Sichote in Latin).

311

Clubiona (Clubiona) stagnatilis Kulczyсskiin Chyzer et Kulczyсski, 1897
MATERIAL. RUSSIA: Belgorod Area: 1 , W of Belgorod, 8 km SE of Kustovoe Village, edge of humid Salix forest, 07.06.1996, leg. et Borisovskiy Distr., Krasivo Village, Vorskla flooded meadow, 18.06.1997, AP; Yakutia: 1 ( alas-pond, 24.07.1977, SK; 1 , 1 (ZMMU), pond, 27.07.1977, SK. 3 (PC), 20 km wet meadow and det. AP; 1 (PC), River floodplain, ZMT), Elgyay, big Toibokhoi, alas-

LITERATURE DATA. RUSSIA. MRM: Kandalaksha Reserve [Mikhailov, 1992b]. KRL [Wo?ny, 1995]. LNN: Nizhnesvirskiy Reserve [Oliger T., 1995, 1996]. MSK [Seifulina, Chernyshev, 2001]. RZN: Okskiy Reserve [Mikhailov, 1992b]. KUR: Tsentralno-Chernozyomniy Reserve [Polchaninova, 2001]. ChVSh [Sidorenko, Dunin, 1997]. ARKh: Pinezhskiy Reserve [Krasnobaev, 1990]. KRV [Oliger M., 1999]. PRM [Esyunin, Efimik, 1996]. EKT [Esyunin, Efimik, 1996]. ChLB [Esyunin, Efimik, 1996]: Troitskiy Reserve [Esyunin, Pakhorukov, 1992], Ilmenskiy Reserve [Polyanin, Lagunov, 1992]. KRG [Mikhailov, 1992b]. TMN: S-Yamal [Korobeinikov, Esyunin, 1995]. NVS [Mikhailov, 1992b; Gorlov et al., 1995; Gorlova, 1999]. KMR [Romanenko, 1998]. KRN: environs of Mirnoe [Rybalov et al., 2002]. Tuva [Mikhailov, 1992b; Logunov et al., 1998-ld; Marusik et al., 2000]. Transbaikalia [Danilov, 1999-ld]. BUR: Baikal Lake: Bolshoi Ushkaniy Isl. [Izmailova, 1989a]. ESTONIA [Vilbaste, 1993]. LITHUANIA [Vilkas, 1992-ld; Relys, 1994-ld; Relys, Dapkus, 2002]. BELARUS: Pripyatskiy National Park [Zhukovets, 1997]. UKRAINE. Rovno [Mikhailov, 1992b]. DNTs [Rudnikova, 1994]. Polesye [Evtushenko, 1991b, 1993a,b]. GEORGIA [Mkheidze, 1997: C. stagnalis]. KAZAKHSTAN. VKO [Mikhailov, 1992b]. KIRGHIZIA [Mikhailov, 1992b; Zonshtein et al., 1996-ld]. DISTRIBUTION. Russia European part, Caucasus, W- & S-Siberia. Baltic countries, Belarus, Ukraine, Georgia, Kazakhstan, Kirghizia. S-, C- & N-Europe, Afghanistan.

skiy Reserve [Esyunin, 1996]. NVS [Mikhailov, 1992b]. KMR [Romanenko, 1998]. KRN: Evenkiya [Mikhailov, 1992b]; environs of Mirnoe [Marusik et al., 2002b; Rybalov et al., 2002]. Tuva [Mikhailov, 1992b; Logunov et al., 1998; Marusik et al., 2000]. Transbaikalia [Danilov, 1999-ld]. BUR: Dzherghinskiy Reserve [Danilov, 1995]. ChIT: Sokhondo Reserve [Mikhailov, 1992b]. ESTONIA [Vilbaste, 1993]. LITHUANIA [Pupiska, 1939: as C. erratica C. Koch; Vilkas, 1992-ld; Relys, 1994-ld; Relys, Dapkus, 2002]. BELARUS. Berezinskiy Reserve [Shavanova, 1995]. UKRAINE. Polesye [Evtushenko, 1991b, 1993a]. DISTRIBUTION. Russia European part, Urals, W- & S-Siberia. Baltic countries, Belarus, Ukraine. S-, C- & N-Europe. Mongolia.

T

HE CAERULESCENS - GROUP

Clubiona (Clubiona) caerulescens L. Koch, 1867
MATERIAL. RUSSIA: 3 (PC), Belgorod Area, 25 km W of Belgorod, environs of Kustovoe Village, 1011.06.1996, leg. et det. AP; 2 (OC, ZMMU), Maritime Prov., Lazovskiy Distr. and Reserve, 1976, 1977, TO. GEORGIA: 1 (ZMMU), 40 km W of Mestia, Kherkhvashi E of Nakra (=Naki), 12501700 m, Quercus, Fagus, Carpinus, Picea, Abies, etc. forest, litter & bark, 21.0821.09.1986, SG; 1 (ZMMU), Algeti State Reserve W of Manglisi, Fagus, Picea, Acer etc. forest, 14001450 m, litter & under bark, 1618.05.1987, SG & KE. AZERBAIJAN: 3 (ZMMU), SW of Kuba, 750 m, Fagus, Quercus, Carpinus etc. forest, litter & under bark, 23.04.1987, SG & KE. ARMENIA: 1 (ZMMU), W of Shamshadyn, halfway btw. Idjevan & Berd, 15001600 m, Fagus, Carpinus, Acer etc. forest, litter & under bark, 2627.05.1987, SG & KE.

Clubiona (Clubiona) subsultans Thorell, 1875
MATERIAL. 1 (SZM), Chelyabinsk Area, Turgask Lake, Betula-Pinus forest, 30.08.1979, leg. Polchanova, det. S.Yu. Rakov; Tyumen Area: 3 (ZMMU), confluence of Ob and Irtysh rivers, mixed forest, under bark, 20.08.1983, AZ: 1 (ZMMU), Ob River, 120 km downstream of Aleksandrovskoe, mixed forest, 23.08.1983, AZ; 2 (ZMMU), Yakutia, Tolbokhoi, 28.07.1977, SK.

LITERATURE DATA. RUSSIA. KRL [Uzenbaev, 1992; Wo?ny, 1995]: Kivach Reserve [Tsellarius, 1993]. LNN [Sokolov, 1960, 1962]: Nizhnesvirskiy Reserve [Oliger T., 1996]. NVG [Esyunin et al., 1993]. TVR: Tsentralno-Lesnoi Reserve [Zheltukhina, 1995]. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993; Matveev et al., 1995]. ChVSh [Sidorenko, Dunin, 1997]. TTR: Volzhsko-Kamskiy Reserve [Aleynikova et al., 1979]. SMR [Krasnobaev, Matveev, 1993]. KRS: Caucasian Reserve [Mikhailov, 1992b]. ARKh [Mikhailov, 1992b]. PRM [Esyunin, 1995; Pakhorukov et al., 1995; Esyunin, Efimik, 1996]. BShK [Efimik, 1997]: Bashkirian Reserve [Ghirfanova et al., 1992; Efimik, 1995a]; Iremel Mt. Ridge [Esyunin, Polyanin, 1990]. ChLB [Esyunin, Efimik, 1996; Efimik, Zolotarev, 1998]. TMN: Yugan-

LITERATURE DATA. RUSSIA. KRL: Kivach Reserve [Tsellarius, 1993]. TVR: Tsentralno-Lesnoi Reserve [Zheltukhina, 1995]. KLZh [Esyunin et al., 1993]. TLS [Esyunin et al., 1993, 1995; Penev et al., 1994]. ORL [Esyunin et al., 1993]. KUR: Tsentralno-Chernozyomniy Reserve [Polchaninova, 2001]. BLG: Les-Na-Vorskle Reserve [Esyunin et al., 1993, 1995; Penev et al., 1994]. VRN [Panteleeva, Karavaev, 1992]. MRS [Matveev, 1991; Krasnobaev, Matveev, 1993; Matveev et al., 1995]. ChVSh [Sidorenko, Dunin, 1997]. ULN [Krasnobaev, 1999]. TTR: Volzhsko-Kamskiy Reserve [Aleynikova et al., 1979]. SMR [Esyunin et al., 1993; Krasnobaev, Matveev, 1993; Dunin, 2001]: Zhiguli Reserve [Esyunin et al., 1993, 1995; Penev et al., 1994; Krasnobaev, 1996]. KOM [Mikhailov, 1992b]. PRM [Esyunin et al., 1993, 1995; Penev et al., 1994; Pakhorukov et al., 1995; Esyunin, Efimik, 1996; Esyunin & Mazura, 2000]. EKT: Visimskiy Reserve [Esyunin & Efimik, 1996; Ukhova, Esyunin, 1996; Ukhova, 2001]. BShK [Ghirfanova et al., 1992; Esyunin, Efimik, 1996; Efimik, 1997]: Shulgan-Tash Reserve [Esyunin et al., 1993; Efimik, Gulyashchikh, 1995]; Bashkirian Reserve [Ghirfanova et al., 1992; Efimik, 1995a]. ChLB [Esyunin, Efimik, 1996]: Troitskiy Reserve [Esyunin, Pakhorukov, 1992]. ORN [Esyunin, Efimik, 1996; Efimik et al., 1997]. TMN: Yuganskiy Reserve [Esyunin, 1996]. KMR [Romanenko, 1998]. ALT [Mikhailov, 1992b; Marusik et al., 1996]. KRN: S part [Mikhailov, 1992b], environs of Mirnoye [Marusiket al., 2002b; Rybalovet al., 2002]. Tuva [Mikhailov, 1992b; Logunov et al., 1998; Marusik et al., 2000]. YaKT: Middle Lena flow [Belimov, 1975]. MGD: environs of Magadan [Marusik et al., 1992-ld]. IRK [Mikhailov, 1992b]. Transbaikalia [Danilov, 1999-ld]. BUR [Mikhailov, 1992b]: Vitimskiy Reserve [Krasnobaev, 1994]; Dzherghinskiy Re-


312

K.G. Mikhailov
Nauka Publ. 91 pp. [in Russian] Apostolov L.G., Onchurov M.V. 1998. [New taxa in araneofauna of Crimea] // Uch. zap. Simferopolsk. gos. univ. Biol., Matem., Psikhol., Fiz. kult., Fiz., Khim. No.5. P.36 [in Russian]. Azarkina G.N. 1999. [Materials on the spider (Arachnida, Aranei) fauna of North-West Altai and its foothills] // Osobo okranyaemye prirodnye territorii Altaiskogo kraya i sopredelnykh regionov, taktika sokhraneniya vidovogo raznoobraziya i genofonda. 4 region. nauch.-prakt. konf. 2729 okt. 1999. Barnaul. P.7375 [in Russian]. Azheganova N.S., Stenchenko T.I. 1977. [To the spider fauna of the Amur Area] // Voprosy arakhnoentomologii. Perm: Perm Univ. P.106111 [in Russian]. Belimov G.T. 1975. [Some data on the arachnofauna of the Middle Lena Valley] // Biologhicheskie resursy Yakutii. Yakutsk: Yakutsk. Univ. P.4243 [in Russian]. Bukhkalo S.P. 1994. [Biotopical distribution of terrestrial invertebrates in the Kontaktoviy Stream Basin] // Kompleksnye ekologhicheskie issledovaniya na statsionare Kontakt. Vladivostok: Dalnauka (for 1993). P.226250 [in Russian]. Bukhkalo S.P. 1995. [Terrestial invertebrate population of Kontakt Station. Part 1. Species composition and biotopical distribution of terrestrial invertebrates in 1987]. Magadan: Inst. of Biological Problems of the North. 36 pp. [in Russian] Bukhkalo S.P. 1997. [Terrestial invertebrate population of Kontakt Station. Part 3. Species composition and biotopical distribution of terrestrial invertebrates]. Magadan: Inst. of Biological Problems of the North. 24 pp. [in Russian] Bukhkalo S.P. 2002. [Species composition of terrestrial invertebrates in poplar-chosenia ecosystems of north-east Russia] / / Russian Entomol. J. Vol.11. No.1. P.101110 [in Russian, with English summary]. Danilov S.N. 1995. [Spiders of the Dzherghinskiy Reserve] // Bioraznoobrazie ekosistem Pribaikalya. Trudy zapoved. Dzherginskiy. Ulan-Ude: Buryatskoe knizhnoe izdatel'stvo. No.1. P.5364 [in Russian]. Danilov S.N. 1997. [New data on the spider fauna of Dzherghinskiy Reserve] // Sostoyanie i problemy okhrany prirodnykh kompleksov Severo-Vostochnogo Pribaikalia. Trudy gos. zapoved. Dzherginskiy. Ulan-Ude. No.2. P.5559 [in Russian], 110 [English summary]. Danilov S.N. 1999. [3.3.3. Spiders of Transbaikalia] // I.Yu. Koropachinskiy, V.M. Korsunov (eds.). Bioraznoobrazie Baikalskoi Sibiri. Novosibirsk: Nauka Publ., Siberian Branch. P.266274 [in Russian]. Dunin P.M. 2001. [3.5. Soil mesofauna: terrestrial arthropods, with insects excluded] // G.S. Rosenberg (ed.). Yabloneviy Ovrag: ekologicheskoe sostoyanie antropogenno narushennoi territorii / Ekologicheskaya bezopasnost i ustoichivoe razvitie Samarskoi oblasti. Togliatti. No.10. P.4558 [in Russian]. Dunin P.M., Mamedov A.A. 1992. [Spiders in cotton fields of the southeastern part of Azerbaijan] // Byull. Moskovsk. obshch. ispytat. prirody. Otdel biol. Vol.97. No.6. P.5361 [in Russian, with English summary]. Efimik V.E. 1995a. [Spiders (Aranei) of the Bashkirian State Reserve] // Fauna i ekologiya paukov. Perm: Perm Univ. (for 1994). P.98116 [in Russian]. Efimik V.E. 1995b. [Zoogeographical analysis of the spider fauna of the western macroslope of South Urals] // Vestnik Permsk. Univ. Perm. No.1. Biologiya. P.102107 [in Russian, with English summary]. Efimik V.E. 1997. [Biotopical arrangement of the spiders of Bashkiria] // Vestnik Permsk. Univ. Perm. No.3. Biologiya. P. 128138 [in Russian, with English summary]. Efimik V.E., Esyunin S.L., Kuznetsov S.F. 1997. Remarks on the Ural spider fauna, 7. New data on the fauna of the Orenburg Area (Arachnida Aranei) // Arthropoda Selecta. Vol.6. No.1/ 2. P.8590. Efimik V.E., Gulyashchikh E.V. 1995. [The spider fauna of pinebroadleaved forests of South Urals] // Fauna i ekologiya paukov. Perm: Perm Univ. (for 1994). P.116131 [in Russian]. Efimik V.E., Zolotarev M.P. 1998. [Additional data on the fauna and ecology of spiders (Arachnida: Aranei) of the Chelya-

serve [Danilov, 1995]. KhAB: Bolshoi Khekhtsir Mt. Ridge [Kim, Kurenshchikov, 1995-ld]. SAKh: Sakhalin [Mikhailov, 1992b; Marusik et al., 1993b-ld]. ESTONIA [Vilbaste, 1993]. LITHUANIA [Pupiska, 1939: C. coerulescens; Vilkas, 1992-ld: C. coerulescens; Relys, 1994-ld]. UKRAINE. KRV [Esyunin et al., 1993]. ODS [Esyunin et al., 1993]. SUM [Esyunin et al., 1993, 1995; Penev et al., 1994]. Polesye [Evtushenko, 1991a,b, 1993a]. The Crimea [Onchurov, 1998: caemlescens]. MOLDAVIA [Esyunin et al., 1993, 1995; Penev et al., 1994]. KAZAKHSTAN. SKO [Mikhailov, 1992b]. DISTRIBUTION. Russia European part, Caucasus, Siberia, Far East (sporadically).Baltic countries, Belarus,Ukraine, Moldavia, Georgia, Azerbaijan, Armenia, N-Kazakhstan. S-, C- & N-Europe, Japan (Hokkaido), China (Inner Mongolia, Jilin).

T

HE CHABAROVI - GROUP

Clubiona (Clubiona) chabarovi Michailov, 1991
Clubiona chabarovi Mikhailov, 1991: 224225, figs. 4449 ().

LITERATURE DATA. RUSSIA. KhAB: S part [Kurenshchikov, 1997b]; Bolshoi Khekhtsir Mt. Ridge [Kurenshchikov, 1997a]. DISTRIBUTION. This species is known only from the type locality.

ACKNOWLEDGEMENTS. I deeply obliged to my colleagues and friends, Dr. Yu.M. Marusik (Magadan, Russia), Dr. D.V. Logunov (Novosibirsk, Russia/Manchester, Great Britain), Dr. S.L. Esyunin (Perm, Russia), Dr. D.K. Kurenshchikov (Bolshekhekhtsyrskiy Reserve, Khabarovsk Province, Russia), Mr. L.A. Trilikauskas (Bureinskiy Reserve, Khabarovsk Province, Russia), Ms. G.N. Azarkina (Novosibirsk, Russia), Dr. T.I. Oliger (Nizhnesvirskiy Reserve, Leningrad Area, Russia), Dr. R.R. Seifulina (Moscow, Russia), Mr. A.V. Ponomarev (Razdorskaya Settlement, Rostov Area, Russia), Dr. A.V. Tanasevitch (Moscow, Russia), Dr. S. Koponen (Turku, Finland), Mr. N.M. Kovbluyk (Simferopol, Ukraine), Dr. E.F. Guseinov (Baku, Azerbaijan) and Mr. A.V. Gromov (Almaty, Kazakhstan) for providing me with numerous specimens of Clubiona. I am grateful to numerous people in Russia and worldwide who provide me with reprints and copies of arachnological papers. Kind assistance from the curators, Prof. Dr. J. Prуszyсski (IZW), Dr. V.A. Krivokhatskiy (ZIN), Dr. M. Saaristo (ZMT) is greatly appreciated. Dr. K. Thaler (Innsbruck, Austria) kindly lent me comparative specimens of C. dvoraki from his personal collection. Many thanks again to Drs D.V. Logunov and Yu.M. Marusik, who kindly reviewed logistics of the ms. Dr David Penney (Manchester, Great Britain) is obliged for his kind linguistic help. The immense patience and understanding of my wife, Elena A. Mikhailova and son Yaroslav K. Mikhailov, during my home work on weekends and during the evening, especially with literature data, made finishing this delayed work possible.

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