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Russian Entomol. J. 14(4): 251-262

ї RUSSIAN ENTOMOLOGICAL JOURNAL, 2005

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae) Заметки о номенклатуре, таксономии и фаунистике ряда видов из рода Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae) H. Fery1, P.N. Petrov2 Х. Фери1, П.Н. Петров2
1 1 2 2

Rдuschstr. 73, D-13509 Berlin, Germany. e-mail: hanfry@aol.com 73, D-13509 , . Moscow South-West Gymnasium 1543, ul. 26 Bakinskikh Komissarov 3-5, 119571 Moscow, Russia. e-mail: bloomsday@newmail.ru - 1543, . 26 3-5, 119571 , .

KEY WORDS: Coleoptera, Dytiscidae, Hydroporus, lectotypes, new synonyms, first records. : Coleoptera, Dytiscidae, Hydroporus, , , . ABSTRACT. Several species of Hydroporus Clairville, 1806, chiefly members of the Hydroporus planus-group, are revised at least in part, and information on the taxonomy and the distribution of these and some other species of the genus is given. The following new synonymies are proposed: Hydroporus transgrediens Gschwendtner, 1923 = Hydroporus discretus ponticus Zaitzev, 1927 syn.n.; Hydroporus goldschmidti Gschwendtner, 1923 = Hydroporus goldschmidti var. recidivus Gschwendtner, 1923 syn.n., and Hydroporus tessellatus (Drapiez, 1819) = Hydroporus habelmanni Wehncke, 1876 syn.n. Lectotypes of the following taxa are designated: Hydroporus corsicus Wehncke, 1872, Hydroporus cyprius Rйgimbart, 1878, Hydroporus discretus ponticus Zaitzev, 1927, Hydroporus distinguendus var. estrellensis Schaufuss, 1882, Hydroporus limbatus Aubй, 1838, Hydroporus brucki Wehncke, 1875, Hydroporus analis Aubй, 1838, Hydroporus decipiens Sharp, 1878, Hydroporus habelmanni Wehncke, 1876, and Hydroporus bicolor G. Mьller, 1933. Several first records are given: H. transgrediens from Armenia, Azerbaijan, Iran, Turkey, and Turkmenistan, H. pubescens (Gyllenhal, 1808) from Georgia and Asian Russia (the Urals), and Hydroporus marginatus (Duftschmid, 1805) from Iran and Kazakhstan. . ( , ) Hydroporus Clairville, 1806, Hydroporus planus. , . : Hydroporus transgrediens Gschwendtner, 1923 = Hydroporus discretus ponticus Za itze v, 1927 s y n.n. ; H y d r opo r u s go l d sc hm i d ti Gschwendtner, 1923 = Hydroporus goldschmidti var. recidivus Gschwendtner, 1923 syn.n. Hydroporus tessellatus (Drapiez, 1819) = Hydroporus habelmanni Wehncke, 1876 syn.n. : Hydroporus corsicus Wehncke, 1872, Hydroporus cyprius Rйgimbart, 1878, Hydroporus discretus ponticus Zaitzev, 1927, Hydroporus distinguendus var. estrellensis Schaufuss, 1882, Hydroporus limbatus Aubй, 1838, Hydroporus brucki Wehncke, 1875, Hydroporus analis Aubй, 1838, Hydroporus decipiens Sharp, 1878, Hydroporus habelmanni Wehncke, 1876 Hydroporus bicolor G. Mьller, 1933. : H. transgrediens -- , , , , H. pubescens (Gyllenhal, 1808) -- () Hydroporus marginatus (Duftschmid, 1805) -- .

Introduction
The present authors have been dealing with members of the genus Hydroporus for several years and have studied a lot of material from a number of museums and private collections. These studies allow a new valuation of some taxa, and we take this opportunity to propose several new synonymies and designate some lectotypes. Further data of interest, such as on the distribution of these and some other taxa, have also been included, as well as several first records.

Material and methods
The following acronyms are used for the collections where material is located: BMNH British Museum of Natural History, London, Great Britain (R. Booth) CHF coll. H. Fery, Berlin, Germany; property of the NMW CJH coll. J. Hбjek, Praha, Czech Republic

252 CJS CGW CKE CLH DEUM

H. Fery, P.N. Petrov
( l e ct o t yp e d es i g n a t i o n o f H. d i s cr et us ) ; Ni l ss o n , 2 00 1 : 1 5 6, 2 0 0 3 : 6 0. MATERIAL. Type material: Lectotype of Hydroporus discretus designated by Balke & Fery [1993: 95], stored in the M N HN. Lectotype of Hydroporus corsicus (by present designation): #, a very small square label without any text, "Haag" [= collector?, hw Wehncke], "Corsica" [hw Wehncke], "Musйum Paris, ex coll. R. Oberthur, ex Wehncke", "Lectotype, Hydroporus corsicus Wehncke, 1872, des. H. Fery 2005" [red, printed]; placed in the drawer below a label "corsicus mihi" [hw Wehncke] (MNHN). Paralectotype: 1 $, "Habelm" [= collector Habelmann, hw Wehncke], "Musйum Paris, ex coll. R. Oberthur, ex Wehncke", and the respective red paralectotype label (MNHN). Notes: The paralectotype is assumed to belong to the syntype series, since it is prepared in the same manner as the lectotype. Wehncke [1872: 163] compared his new taxon with Hydroporus longulus Mulsant & Rey, 1861, a species which does not belong to the planus-group, but to the Hydroporus longulus-group. And, indeed, the two types and some other specimens studied from Corsica are somewhat elongate and at first glance resemble H. longulus. Lectotype of Hydroporus cyprius (by present designation): #, "Chypre" [green, hw?], "cyprius Rйgimb." [hw Rйgimbart], "Musйum Paris, Coll Maurice Rйgimbart, 1908", "Lectotype, Hydroporus cyprius Rйgimbart, 1878, des. H. Fery 2005" [red, printed] (MNHN). Paralectotypes: 7 exs., one specimen with first and second labels as in the lectotype, the rest only with the first label; coll. Sedillot (MNHN). Each paralectotype is provided with its respective red label. Additional material examined: France, Corsica: 2 exs., "Corse", "Musйum Paris, ex coll. R. Oberthur"; 4 exs., "Vizzavona, Corse", "Hydroporus Corsicus ([1]895)", "Corse, Damry", "Musйum Paris, ex coll. R. Oberthur"; 2 exs., "Rouisseau, Zonza", "Hydroporus Corsicus", "Corse, Damry", "Musйum Paris, ex coll. R. Oberthur"; 1 ex., "Corsicus", "Corse, Damry", "Musйum Paris, ex coll. R. Oberthur" (all in MNHN). 1 #, "F 37 (2B) Morosaglia, (Corte), Bocca di Riscamone, 570 m. 09њ15E 42њ26N. 01.10.[19]91. leg. GER [= Gerecke]"; 1 #, "Bastia, Corsika, V.1907, F. Hopp"; 1 $, "10.7.[19]86 Corse, Pinito, Porto, Quellmoos, Fery leg."; Cyprus: 7 exs., "31.3.[19]96 Zypern, nr Kannaviou, N Pafos, Bach, Fery leg."; 3 exs., "10.4.[19]96 Zypern, nr Kannaviou, N Pafos, Ezousa river, Fery leg."; 1 $, "4.4.[19]96 Zypern (Lim.), Mandria, SW Platres, Bach, Fery leg."; 5 exs., "8.4.[19]96 Zypern (La.), Vavatsinia, W Lefkara, Bach, Fery leg." (all in CHF).

coll. J. St'astnэ, Liberec, Czech Republic coll. G. Wewalka, Vienna, Austria coll. Ц.K. Erman, Erzurum, Turkey coll. L. Hendrich, Berlin, Germany Department of Entomology, Moscow Lomonosov State University, Russia (V.Y. Savitsky) IRSN Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium (M. Cludts, K. Desender) MNB Museum fьr Naturkunde, Humboldt-Universitдt, Berlin, Germany (F. Hieke, M. Uhlig, B. Jaeger) MCTR Museo Civico di Storia Naturale, Trieste, Italy (G. Alberti, A. Colla) MNHN Musйum National d'Histoire Naturelle, Paris, France (H. Perrin) MRTO Museo Regionale di Scienze Naturali, Torino, Italy (M. Daccordi) NMB Naturhistorisches Museum Basel, Switzerland (M. Brancucci) NMW Naturhistorisches Museum Wien, Austria, Vienna (M. Jдch) OLML Oberцsterreichisches Landesmuseum, Linz, Austria (F. Gusenleitner, C. Reitstдtter) SMNS Staatliches Museum fьr Naturkunde Stuttgart, Germany (W. Schawaller) ZISP Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia (A.G. Kirejtshuk) ZSM Zoologische Staatssammlung, Mьnchen, Germany (M. Baehr, M. Kьhbandner) The following abbreviations are used in the text: TL (total length), MW (maximum width), hw (handwriting), and hw? (handwriting of an unknown person). For the ink drawings, male genitalia were studied wet. The terminology denoting genitalia orientation follows Miller & Nilsson [2003]. All lectotype designations are made in order to support the stability of the nomenclature (cf. Article 74.7.3 of the ICZN [1999], and Anonymous [2003]). Specifications of localities are taken from standard sources, such as Stiehlers Handatlas (Gotha, 1928/30), Microsoft Encarta World Atlas 2000, and The Times Atlas of the World, Comprehensive Edition (London, 1997). We have adopted the style of the latter source for the presentation of the names and co-ordinates of most localities. The authors' remarks are given in square brackets.

DESCRIPTIVE NOTES. Specimens from Corsica coincide more or less with central European ones. Those from Cyprus often have the elytra somewhat more brownish; in addition, we have observed that the gonocoxosterna are slightly longer and the gonocoxae more rounded apically. With respect to the high variability of H. discretus, however, these observations seem not to justify a treatment as proper species or subspecies. See also the `Discussion' at the end of the section about Hydroporus transgrediens Gschwendtner, 1923. The male genitalia of a specimen from Bavaria, Germany, are given in Figs 1-3.

Results
Hydroporus planus-group Hydroporus discretus Fairmaire & Brisout de Barneville, 1859
Fairmaire & Brisout de Barneville in Fairmaire, 1859: 28 Type locality -- France, Marly = Hydroporus corsicus Wehncke, 1872: 163 (Type locality Fran ce, C ors ica) = Hydroporus cyprius Rйgimbart, 1878: 352 (Type locality C y p ru s) Guignot, 1959: 393 (synonymy of H. corsicus); Miller et 1997: 27 (synonymy of H. cyprius); Balke & Fery, 1993: -- -- al., 95

Hydroporus transgrediens Gschwendtner, 1923
Gschwendtner, 1923: 109 (H. pubescens transgrediens) Type locality -- Transcaspien, Neu-Saratow = Hydroporus discretus ponticus Zaitzev, 1927: syn.n. (Type l o cal it y -- Ar men ia, Y erev an d istr., D ara i ag [ = Tsag h k adzor], ca. 40 km NNE Yerevan) Zi mmermann , 19 31 : 13 7 (p ub es cens t ra ns gr ed iens ); Gui gn ot, 1 932 : 372 (p ub escens var. t ra nsg red iens ); Zai tzev, 19 51 : 56 (pub escens tr ansgr ed ie ns ), 195 3: 165 (pub escens tr ansgr ed iens), 169 (discretus ponticus), 1972: 174 (pubescens transgrediens), 179 (discretus ponticus); Wewalka, 1992: 55 (lectotype des ignat ion o f H. pubes cens t ransg rediens); Бdбm, 19 96: 59 (ponticus); Nilsson, 2001: 156 (discretus ponticus), 158; Nilsson, 2003: 61 (discretus [= ponticus]), 65.

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae)

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Figs 1-12. Male genitalia of Hydroporus spp.: 1-3 -- H. planus; 1-2, 4-5, 7-8, 10-11 -- penis; 3, 6, 9, 12 -- vie w. Рис. 1-12. Гениталии самцов Hydroporus spp.: 1-3 12 -- H. planus; 1-2, 4-5, 7-8, 10-11 -- пенис; 3, 6, 9

H. discretus; 4-6 -- H. transgrediens; 7-9 -- H. goldschmidti; 10-12 -- paramere; 1, 4, 7, 10 -- ventral view; 2-3, 5-6, 8-9, 11-12 -- lateral -- H. discretus; 4-6 -- H. transgrediens; 7-9 -- H. goldschmidti; 10- , 12 -- парамера; 1, 4, 7, 10 -- снизу; 2-3, 5-6, 8-9, 11-12 -- сбоку. gical Institute, Russian Academy of Sciences, St. Petersburg" [yellow], "Lectotype, Hydroporus discretus ponticus Zaitzev, 1927, des. H. Fery 2005" [red, printed] (ZISP). Paralectotypes: Armenia: 14 exs., the same label data as the lectotype; two specimens considerably damaged by dermestids; Georgia: 3 ##, 1 $, "Baraleti [ca. 41.32N 43.29E], prov. Achalk., VI-VII. [19]16."; 1 ex., "Ordzhoch [not found on any map], doroga [= road], 28.6.[19]09, Bat. ob. [= Batumi oblast]" [Cyrillic], considerably damaged by dermestids; Turkey: 1 #, "Bajburt vic., distr. E r ze r u m , 1 8. VII . [19] 1 7" ; 1 $ , " Ke s ala r , d ist r . E r z e r um, 29.VII.[19]17", somewhat damaged; 1 ex., "Sadach, 7.VII.[19]17, E...itter [? illegible]" [hw?], considerably damaged by dermestids; according to Zaitzev [(1927: 17)] this locality is in "Trapezunt [= Trabzon] Province"; Ukraine: 3 exs., "Krim. Jalta district, Tauschan-Basar, 15.6.[19]07. W. Pliginskiy" [Cyrillic], one specimen considerably damaged by dermestids; 1 $, "Krim. Jalta district, Tauschan-Basar, 23.6.[19]07. W. Pliginskiy" [Cyrillic]; 1 #, "Krim. Angarsk-pass, puddle near road, 27.6.[19]07. W. Pliginskiy" [Cyrillic]; 1 ex. "Krim", considerably damaged by dermestids; 1 #, "Sebastopol, Krim 22.3.[19]06, W. Pliginskiy"; 1 $, "Sebastopol, Krim 8.4.[19]06, W. Pliginskiy"; 1 $, "Sebastopol, Krim 26.V.[19]06, W. Pliginskiy"; 1 $, "Tauria 89, Aluschta [ca. 44.41N 34.31E]" (ZISP). 1 #, "Sebastopol, Krim, Inkerman [suburb of Sevastopol], 6.3.[19]10 W. Pliginskiy." [Cyrillic in part], "334", "Hydroporus sp. n." [hw Reitter], "Reitter det." [printed], "H. planus var. aut n. sp., V. Pliginski. coll." [hw Reitter in part], "Reitter det." [printed], "Hydroporus discretus ponticus

MATERIAL. Type material: Lectotype of Hydroporus pubescens transgrediens: #, "Transcaspia., Neu-Saratow"* [printed], "Type, Gschw" [greyish brown label, hw Gschwendtner], "Coll. Gschwendtner" [printed], "Hydroporus pubescens transgrediens Gschw" [hw Gschwendtner?], "Type" [red, hw?], "Lectotypus, Hydroporus transgrediens Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML). Paralectotype: 1 #, first three labels as in the lectotype, "pubescens v. transgrediens Gsch., det. Gschwendtner" [hw Gschwendtner], "Type" [red, hw?], "Paralectotypus, Hydroporus transgrediens, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML). Lectotype of Hydroporus discretus ponticus (by present designation): #, "DaraИiИag [= Darachichag = Tsaghkadzor, ca. 40.32N 44.45E] & vic, distr. Erivan, 24.VIII.[19]12.", "Zoolo* We have not found a locality named "Neu-Saratow" [= New S arato v = N o vyy Sa r at o v] o n a ny ma p . T h ere ex i s ts t he v illa g e N ovos arat ovk a [ w h i c h ca n b e r ough l y t ra ns lated from Russian as "New Saratov Village"] (40.36N 45.35E) in Aze rb aija n , a c oun t r y t h at, ho we v e r , do es n o t b el o n g to "Transcaspia". Possibly, "Transkaspien [= Transcapia]" is a mi s spell i ng o f " Tra ns cau c asia" , and Novo s arato vk a mi ght be the correct type locality, a solution which we assume to be the most probable. Nilsson [2003: 65] provides as distribution area "TM" [= Turkmenistan], which is certainly due to t h e st a te m e n t i n W e w al k a [ 1 9 92 : 5 4 ] w h o g a v e "T urkmeniya" as distribution area.

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H. Fery, P.N. Petrov
emerak, 23.09.2001, Erman, Ц.K. leg." (CKE); 7 exs., "18.9.2000 (TR) prov. Erzurum, Yedigцller, nr Uzunkavak, ca. 3000 m, ca. 20 km E spir, Ц.K. Erman leg." (CHF); 9 exs., "18.4.2000 (TR) Erzurum, ca. 28 km SW Erzurum, N pass, ca. 1800 m, ponds on meadow, Fery leg." (CHF); 3 exs., "18.4.2000 (TR) Erzurum, ca. 43 km SW Erzurum, S pass, ca. 2000 m, ponds on meadow, Fery leg." (CHF); 1 ex., "Turkey 20.VI.2003, Erzurum vil. [= province] (ca 15 km W), Ilica env. (pools; sweeping), (39њ55'N, 41њ06'E; 1770 m), Jiшн Hбjek & Josef Hotovэ leg." (CJH); 1 ex., "TR 5.6. 1987, Van - Baskale 2600 m, Gьzeldere P. [= pass], leg. Jдch (62)" (NMW); 2 exs., "Turkey 28.-29.VI.2003, Van vil. [= province] (ca 40 km SW), Geva? env. (stream), (38њ16'N, 43њ03'E; 1880 m), Jiшн Hбjek & Josef Hotovэ leg." (CJH); 2 exs., "Turkey 27.-28.VI.2003, Van vil. [= province] (ca 45 km SE), Guselsu - Ho?ap (stream), (38њ18'N, 43њ47'E; 2070 m), Jiшн Hбjek & Josef Hotovэ leg." (CJH); 1 $, "Umg. [= environs] Kayabo az, se Geva?, 2100-2200 m, 31.VII.1985"*, "SьdostAnatolien, Heinz leg." (NMB); 5 exs., "Turkey 17.-18.VI.2003, K. [= Karaman] Mara? (ca 25 km SW), Fatih env. (stream), (37њ26'N, 36њ41'E; 1115 m), Jiшн Hбjek & Josef Hotovэ leg." (CJH); 2 exs., "TR - Isparta, Gelendost, 9-VI-[19]81 G. Sama" (CHF); 1 #, "TR - Antalya 24.5.[19]91, n Altinyaka 1100 m, leg. Schцdl (32)" (NMW); 1 ex., "Turkei 21.6.[19]97, Gцreme, lgt Jiшн Hбjek", "Hydroporus discretus ponticus Zaitzev, Jiшн Hбjek det. 2001" (CJH). 5 exs., "Simav Dag [ca. 39.07N 28.30E], 1978, 100/1500 m 16.VIII.", "W Anatolien, leg. Casale" (NMB); 3 exs., "TR 15.6. 1987, 80 km w. Istanbul, leg. M. Jдch (79)" (NMW). Iran (first record): 1 #, "Danavar-dagh, s.e. [= southeast of] Ushnuiyeh, 1600 m, 10.VIII.1969", "Azerbaijan (Iran), Heinz leg." (NMB). 21 exs., "15.8.1998 Iran, Kohkiluyeh & Boyer Ahmad, 7 km SW Yasuj, Elmi & Fery leg.", "Cheshmeh Sare-Ab Taveh, brook and pools (with stagnant water) (#2106)"; 4 exs., "15.8.1998 Iran, Fars, 17 km SE Sepidan (= Ardakan), Shesh Pir", "ditch with running water, Elmi & Fery leg. (#2108)"; 10 exs., "18.4.2001 Iran, Fars, 17 km SE Sepidan (= Ardakan), Shesh Pir", "ditch with running water, Elmi & Fery leg."; 168 exs., "16.8.1998 Iran, Fars prov., Bamoo-Nat. Park, spring Cheshmeh-ye-Ghanbari, Elmi & Fery leg. (#2109)"; 9 exs., "16.8.1998 Iran, Fars, Nat. P. Bamoo, Darreh-Bishen spring-stream system, Elmi & Fery leg. (#2113)" (all in CHF). Turkmenistan (first record, except that from the doubtful locus typicus -- see above): 1 $, "Turkmenia, Bolshoi [= big] Balkhan [mountain range, ca. 200 km E Krasnovodsk, ca. 40N 55E], 10.VII.[19]40 leg. Starostin"; 1 #, "Turkmenia, Mary st a t io n [ca . 3 7 . 3 4N 6 1 . 4 8E; M a r y = a nc ie nt Me r w ], l eg Starostin" [Cyrillic]; 1 #, "Aj-Dere, W Kopet-dag, Iljitshev [collector], 30.IV.[1]952"; 1 #, "Kara-kala e nv., Turkmen., Steinberg [collector] 26.IV.[1]953" [Cyrillic] (all in ZISP). 1 #, 3 $$, "Turkmenia, 20.05.[19]87., mountain range Kopet Dag, Schlucht [= gorge] Kuru Chudan, Wolkow D.A. leg." (DEUM). Three of the specimens from Turkmenistan are with rather yellowish spots near the base of elytra.

Zaitz., Shaverdo H. det. 2000"; 1 $, "Krim, Jalta Kreis, village [?] Saki., 8.VIII.[19]13[?] W. Pliginskiy" [Cyrillic in part], "Hydroporus discretus ponticus Zaitz., Shaverdo H. det. 2000"; 1 #, "Tauria, village Saki, Yepatoria distr., 7.8.[19]12 W. Pliginski" [Cyrillic in part], "H. planus var. aut n. sp., V. Pliginski. coll." [hw Reitter in part], "Reitter det." [printed], "Hydroporus discretus ponticus Zaitz., Shaverdo H. det. 2000"; 1 $, "Tauria, village Saki, Yepatoria distr., 26.8.[19]12 W. Pliginski" [Cyrillic in part], "H. planus var. aut n. sp.?" [hw Reitter], "Reitter det." [printed], "Hydroporus discretus ponticus Zaitz., Shaverdo H. det. 2000"; 1 #, "Saki, Krim 18.8.[19]10, W. Pliginski" [Cyrillic in part], "Hydroporus nigrita" [hw Reitter], "Reitter det." [printed], "T.k. nadkrylya bez shagrenirovannogo fona, to eto discretus, v spisok voshyol kak H. nigrita, N 388 yeshcho raz poslat na proverku, W. Pliguinskii [= Since the elytra are without reticulate (shagreen) surface, this is discretus, was included in the checklist as H. nigrita, send No. 388 to be checked again]" ["W. Pliguinskii" printed in Latin; rest in Cyrillic, hw Pliginski?]. Unknown localities: 1 ex., "st. Passanaur, dist. Duset, VII.[19]13", sp ec imen a lmost to tal ly d estr oy ed b y d er me st id s; 2 e xs., "Suram, 6.VI.[19]17, Ahnger", one specimen considerably damaged by dermestids (all in ZISP). Each paralectotype is provided with its respective red label and the yellow label of the Zoological Institute of the Russian Academy of Sciences. Additional material examined: Ukraine: 2 ##, "Tauria [= Crimea]" [blue label]; 1 ex., "Tauria 89, Mshatka [= Yuzhnoye, ca. 44.24N 33.48E]" [hw?], considerably damaged by dermestids (all in ZISP); 10 exs., "Crimea (Ukraine), env. Morskoje [ca. 44.50N 34.48E], 24.04.1994" (NMB, CHF). Russia: 10 exs., "Kaukasus, Dagestan, Kurush, 2200-2400 m, 3.-6.7.1991, Ma rt ens, Sc ha wa ll er et a l." ( SMNS, C HF). 1 #, "USSR, Caucasus, Krasnaja Poljana, 4.-8.7.1988, Dr. L. Danйk lgt." (CJS). Georgia: 2 exs., "Kaukasus, Tiflis", one specimen considerably damaged by dermestids (ZISP). Armenia (formally a first record under the name H. transgrediens): We have studied a large amount of specimens collected by Shaverdo and Schillhammer in May 2001. The respective records are listed in Shaverdo [2003] under the name H. discretus ponticus and shall not be repeated here. Azerbaijan (first record): 6 exs., "Disar nr O r d u bad , N a c hitsh e w a ns ki r e gion , Zno jko [co ll e ctor] 22.VII.[1]933", "Hydroporus discretus ponticus Zaitz., Shaverdo H. det. 2000" (ZISP); Turkey (formally a first record from Turkey under the name H. transgrediens, however known from this country under the name H. discretus ponticus since Zaitzev [1927]): 4 exs., "Turquie: Artvin, SW Artvin, 1900 m, 9.VI.1986", "Besuchet - Lцbl, Burckhardt" (CHF); 3 exs., "TR - Artvin (47), Artvin - Savsac, Schцdl 4.6.[19]89" (CHF, further specimens in NMW); 1 #, "Turkey, Artvin prov., 8 km E av?at, Karagцl Sahara Nat. Park, 1930-2170 m, 4.-5.VII.2004, Jiшн Hбjek & Jan Rщ iИka leg.", "41њ13.5'N 042њ27.1'E - 41њ13.5'N 042њ28.0'E, pasture / wet coniferous forest, (dominant Picea, Abies)" (CJH). 1 $, "ca. 13 km S Posof [ca. 41.30N 42.41E, in Ardahan prov.], 2000 m 18.VII.1983", "Anatolien, W. Heinz" (NMB). 1 #, "TR - Rize, kisdere, Anzer Kцyь Yaylasi, Gцllerin Alt ndaki Dere, Brikinti, 29.6.[20]01, Erman, Ц.K. leg." (CKE). 1 #, 2 $$, "TR - Rize, kisdere, Sivrikaya Kцyь, Kцprь, Yan ndaki Il ca, 29.6.2001, Erman, Ц.K. leg."; specimens rather small, TL ca. 3.3 mm (CKE, CHF). 2 exs., "Turcia 3.IV.2000, Bolu vil. [= province], 10 km E Gerede, J. Hбjek & M. Mikбt leg.", "Hydroporus discretus ponticus Zaitzev, Jiшн Hбjek det. 2001" (CJH); 1 #, 1 $, "18.7.1992 TR Erzurum, MT Palandцken, stream on road Erzurum - Зam, 2300 m, Toledo leg." (CHF); 1 #, "21.7.1997 (TR) prov. Erzurum, Tortum - Uzundere road, ca. 5 km N Tortum, artesian well under bridge, Ц.K. Erman leg." (CHF); 6 exs., "26.4.2000 (TR) Erzurum, ca. 10 km S Tortum, ca. 2 km N Gьzelyayla pass, ponds on meadow, Fery leg." (CHF); 8 exs., "26.4.2000 (TR) Erzurum, ca. 14 km S Tortum, ca. 2 km S Gьzelyayla pass, ponds on meadow, Fery leg." (CHF); 1 #, 2 $$, "TR - Erzurum, Oltu, Gьryaprak Kцyь Gцlleri, 22.9.2001, Erman, Ц.K. Leg." (CKE, CHF); 1 #, 1 $, "TR - Erzurum, Oltu, Зarzof Kцyь Gцlleri, 22.9.2001, Erman, Ц.K. Leg." (CKE); 1 #, 2 $$, "TR - Erzurum, Oltu, Gцзedere, 22.9.2001, Erman, Ц.K. Leg." (CKE); 1 $, "TR - Erzurum, enkaya, Turnab Kцyь,

DESCRIPTIVE NOTES. For detailed descriptions of H. transgrediens see Gschwendtner [1923: 109] and Wewalka [1992: 55]. The species externally is similar to Hydroporus pubescens (Gyllenhal, 1808); it has, however, the last abdominal sternum distinctly reticulated, while that of H. pubescens is smooth. It is equally similar to H. discretus, however, on average slightly longer (TL 3.4-4.0 mm; H. discretus: TL 3.25-3.5 mm, reaching 3.8 mm in some southern European specimens), and the colour of the upper surface is dark brown and not black as it is at least in central and northern European populations of H. discretus. In most specimens, the elytra are lighter near the base; in a few specimens from Iran and Turkmenistan we have even observed distinct yellowish spots. The male genitalia are given in Figs 4-6. In lateral view, the penis is slightly more curved in apical third and lacks the convexity of the corrugated membrane in the
* This locality is either located in Siirt Province (ca. 37.53N 41.59E) or in Van Province (ca. 38.09N 43.09E).

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae)
middle part of the ventral surface, which is so characteristic of H. discretus (cf. Fig. 2). DISCUSSION. Although H. transgrediens can be distinguished from "normal" central European H. discretus by its larger total length and differences in the shape of the penis, we cannot exclude the possibility that future investigations will reveal that the species belongs to a difficult complex of species, varieties and forms around H. discretus (see, e.g., Nilsson [2001: 156]), which is distributed over an enormous area reaching from the Canaries, North Africa; most of Europe; large territories in Russia; the Near, Middle, and Far East; and China. Not only the two species and the varieties and forms mentioned above must be included into such studies, but also other taxa which at present are treated as valid species, such as Hydroporus errans Sharp, 1882 and Hydroporus feryi Wewalka, 1992.

255

Hydroporus goldschmidti Gschwendtner, 1923
Gschwendtner, 1923: 101 Type locality -- Kyrgyzstan, Issyk-Kul, Ton [= Tong] river* = Hydroporus goldschmidti var. recidivus Gschwendtner, 1923: 103 sy n.n. (Type locali ty -- Ch ina, Xin jian, Tian Shan, b etween Aksu and Musart pass**) Zi mmer ma nn , 1931 : 13 0, 13 1 ( go l ds c h m i d ti v ar. re ci d i vus ); Z a i t z e v, 1 9 5 3 : 1 6 4 ( i n c l u d i n g g o l d s c h m i d ti v ar . r e c i d i v us ), 1972: 173 (including goldschmidti var. recidivus); V.B. Gueorg u ie v , 19 6 3 : 2 1 6 (a s p o s s i b l e s y n o n y m o f H . g l a s u n o vi Zai tzev, 1905); Wewalka, 1992: 54 (lecto type design ation of H. goldschmidti), 58 (lectotype designation of H. goldschmidti var. recidivus); Nilsson, 1995: 53, 2001: 157, 158 (recidivus), 2003: 61, 64 (recidivus). MATERIAL. Type material: Lectotype of Hydroporus goldschmidti: #, "Jssyk - Kul, Ton-Fluss." [printed], "Type, Gschw" [greyish brown, hw Gschwendtner], "Coll. Gschwendtne r" [pr inted ], "Hydroporu s Gold sc hmi dti # Gsc hw." [hw Gschwendtner], "Type" [red, hw?], "Lectotypus #, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML). Paralectotypes: Kyrgyzstan: 1 $ (strongly matt), "Jssyk - Kul, Ton-Fluss." [printed], "Type, Gschw" [greyish brown, hw Gschwendtner], "Coll. Gschwendtner" [printed], "Hydroporus Goldschmidti $ Gschw." [hw Gschwendtner], "Type" [red, hw?], "Paralectotypus $, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka]; 1 $ (weakly matt), "Jssyk - kul, Ton-FluЯ." [hw Gschwendtner], "Type, Gschw" [greyish brown, hw Gschwendtner], "Coll. Gschwendtner" [printed], "Hydroporus Goldschmidti $ intersex. Gschw." [hw Gschwendtner], "Type" [red, hw?], "Paralectotypus $, Hydroporus goldschmidti Gschw., det. G. Wewalka [19]91" [red, hw Wewalka] (all in OLML). China: 1 #, "Ost-Turkestan, BagratschKul" [printed], "Cotype, Gschw" [greyish brown, hw Gschwendtner], "Hydroporus Goldschmidti Gschw." [hw Gschwendtner], "Type" [red, hw?], "Paralectotypus #, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML); 1 $ (strongly matt), "Ost-Turkestan, Bagratsch-Kul" [printed], "Type!" [hw Zimmermann], "Samml. A. Zimmermann", "Cotypus, Hydroporus Goldschmidti Gschw., Zool. Staatssammlung Mьnchen" [red, hw? non-authorised curatorial designation], "Paralectotype, Hydroporus goldschmidti Gschwendtner, 1923, Fery 2004" [red] (ZSM). Notes: Judging by the original description, Gschwendtner did not know females of this species with smooth elytra. Most probably, he used the term "intersex" in the meaning of "reticulation of the upper surface between that of strongly reticulated females and that of smooth males". 1 #, "Issyk-Kul, Ton-Fluss." [hw?], "Type, * A cc o rd i n g t o S ch ь tze & Kle i nf river rises ca. at 41.52N 77.02E shores of the Issyk-Kul near 42. * *This locality should have about 4 2. 3 0 N 8 0 .4 0E eld an d 11N the [1 9 97 : 14 3], t h e To ng flows into the southern 76.55E following co-ordinates:

Gschw" [greyish brown, hw Gschwendtner], "Coll. Gschwendtner" [printed], "Goldschmidti Gschw., det. Gschwendtner" [hw Gschwendtner?], "Type" [red, hw?], "Paralectotypus #, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML); 1 $ (strongly matt), "Issyk - kul, Ton-Fluss." [hw?], "Type, Gschw" [greyish brown, hw Gschwendtner], "Coll. Gschwendtner" [printed], "Goldschmidti Gschw., det. Gschwendtner" [hw Gschwendtner?], "Type" [red, hw?], "Paralectotypus #, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML); 2 $$ (weakly matt), "Issyk - kul, TonFluЯ." [hw Gschwendtner], "Cotype, Gschw" [greyish brown, hw Gschwendtner], "Paralectotypus $, Hydroporus goldschmidti Gschw., det. G. Wewalka [19]91" [red, hw Wewalka]; one specimen with additional "7211, Goldschmidti" [hw?] (CGW). Zimmermann [1931: 131] studied at least a part of the syntype series. It is assumed that he kept this specimen for his c ol le cti on. We wal ka [199 2: 5 4] ga ve f or "Bagra tsc h-Kul": "China, Sinkiang-Uighur, Bagratsch-Kul or Po-ssu-teng Hu". Today this lake is called "Bosten Hu"; it is in Xinjian province, its co-ordinates are ca. 42N 87E [H. Schьtze in litt.]. Lectotype of Hydroporus goldschmidti var. recidivus: $, abdomen without inner parts, "Thian shan, Aksu-Musart, coll. Winkler" [printed], "Type, Gschw" [greyish brown, hw Gschwendtner], "Goldschmidti var recidivus Gschw" [hw Gschwendtner], "Coll. Gschwendtner" [printed], "Type" [red, hw?], "Lectotypus $, Hydroporus recidivus Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (OLML). Paralectotypes: 1 $, abdomen absent, first two labels as in the lectotype except "Cotype" instead of "Type", "Hydroporus Goldschmidti var. recidivus Gschw" [hw Gschwendtner], "Paralectotypus $, Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91" [red, hw Wewalka] (CGW); 1 #, first two and fourth label as in the lectotype, third label as in t h e f e ma le pa r al e ct o ty p e , " H y d ro po ru s ? g o ld s c h mi d ti Gschw, det. G. Wewalka [19]91" [hw Wewalka], "Paralectotype, Hydroporus recidivus Gschwendtner, 1923, Fery 2000" [red] (OLML). Notes: This male became automatically a paralectotype when Wewalka [1992: 58] selected the lectotype from the syntype series. This is why we have provided this specimen with the respective red label, although Wewalka refrained to do so, because he believed -- correctly -- this specimen to belong to H. goldschmidti, not to H. recidivus. Additional material examined: Uzbekistan: 1 #, 2 $$, "Uzbekistan, Tashkent Province, Yakka-tut near Burchmulla, 26.7.-3.8.2004, 41њ38'N 70њ03'E, Hendrich leg. Loc. 1a"; see Hendrich & Hendrich [2005: 430] (CLH). Kyrgyzstan: 1 $, "Kirghizia: Issyk - Kul bank S Chon-Urjukty [ca. 42.43N 77.50E], 19.-23.VI.1993 1600 m, leg. Schawaller"; 1 $, "Kirghizia: Chatkalskij Alatau, Sary-Celek [near Arkit, ca. 41.47N 71.56E], 27.- 31.V.1993, 1400-1600 m, leg. Schawaller" (all in SMNS). 2 ##, 3 $$ (all smooth), "Kyrgyzstan: Sandalasch G.K. [= Gebirgs-Kette = mountain range], Chatkal riv., ca. 2000 m, 5.VII.1997, leg. Dolin (K - 9/97)" [ca. 41.50N 70.55E, Dolin in litt.]; 3 ##, 6 $$ (all smooth), "Kyrgyzstan: Sandalasch G.K. [= mountain range], Chapchama Pass env., 1600 m, 6.VII.1997, leg. Dolin (K - 10/97)" [ca. 41.40N 70.50E, Dolin in litt.]; most probably the Chap-Chyma Pass, ca. 41.32N 70.48E [see Schьtze & Kleinfeld 1997: 94]; 1 $, "Kyrgyzstan, Ferganski range, nr. Bergut pass [not found on any map], 2015 m, 24.6.1996, leg. Dolin (n 37)"; specimen with pronotum totally reticulated, but elytra smooth (all in NMW); 4 ##, 3 $$, "Kyrgyzstan, W Tien Shan, Chatkal gorge, Stab [not found on any map] env., 29.VII.1998, 1800 m, leg. Dolin (K - 9/ 98)" (NMW); 3 ##, "Kyrgyzstan, Tshatkal valley, Shtab 2000 m, 28.07.[19]99, V. Dolin" (CJH); 2 ##, "Kyrgyzstan, Mt. Rg., Tshatkal, 2000, Shtab, 28\2906[19]98" (CJS); 13 ##, 1 $ (smooth), "28.7.1999 Kyrgyzstan, Tschatkal mountain range, nr Stab, Bach und Pfьtzen, Dolin leg."; found together with Hydroporus glasunovi Zaitzev, 1905 (NMW). China: 1 $, "Thian-Shan, Musart" [printed], "Hydroporus goldschmidti var. recidivus Gschw., det. G. Wewalka [19]72" [hw Wewalka], "Hydroporus recidivus Gschw, det. G. Wewalka [19]91" [hw Wewalka]; 2 ##, 5 $$ (all $$ matt), "Bagratsch-Kul" [some labels printed, others hw Wewalka], the males with "Hydroporus # goldschmidti Gschw, det. G. Wewalka [19]91" [hw Wewalka], the females with similar

256

H. Fery, P.N. Petrov
6.[18]67 " [hw Scha ufuss], "C oll. L.W. SchaufuЯ" [printe d], "distinguend. var estrellensis, Estrella Schauf." [hw Schaufuss], "Zool. Mus. Berlin", "Le ctot ype, Hydroporus dis ting uend us var. estrellensis Schaufuss, 1882, des. H. Fery 2005" [red, p r int e d] ( M N B ) . P a ral e ctot yp es : 1 $ , t h e s p ec im e n is provided with copies of lectotype labels; 1 $, "Estrella, 18/ 667" [hw Schaufuss], "Coll. L.W. SchaufuЯ" [printed], "Zool. Mu s. Be rl i n"; 1 # , " T yp e " [ p r int ed ] , "H y dr . d i s t i ng . var estrellensis Schauf, 18/6 1867 Estrella" [hw?], "Zool. Mus. Berlin"; 1 #, "Type" [printed], "Hydr. disting. var estrellensis, Schauf. I. 18/6 1867 Estrella" [hw?], "Zool. Mus. Berlin" (all in MNB). Each paralectotype is provided with its respective r ed l a bel. No tes : Lectotype a n d th e fir s t paralectotype originally were glued onto the same card; they have been separated by us and each glued onto its own card. The lectotype lacks both hind tarsi and the last two right mid t a r s o m e r es. Addi tion al ma terial examined: Latvia: 1 $, " Kurland, S und ur , Aug. 19 17 , le g. Salc he rt" ( MN B). Geo rgia ( f i rst reco rd) : 7 e xs. , " SSSR-Ca uca sus, Sno [ca. 9 0 k m N Tb il isi, c a . 4 2 .35 N 44 . 38E ] 1 8 0 0 m , 26 . 6.[ 1 9]5 7 Ш нh a " (C H F). Tu rkey : 10 e xs. , " Ul uda g, 185 0 m, 2. VIII. 1984" , " Tьrkey [ sic!] , R. Kinzelbac h et al." , "84 /23" (N MB, CH F). Speci me ns with rat he r l ight elytra ; ri m of pronot um in mo st speci me ns bro wnish. Azerb ai jan : 13 0 e xs. , " 17.5. 1999 Aze rba jdz ha n: Taly sc h [= Taly sh, mo untain s in so uthern Aze rbai ja n], H irkan-Wald , Aurora Umg. [= e nviron s], 6 00 m, Dolin le g."; 2 ##, 2 $$, " 17.5. 1999 Aze rba jdz ha n: Taly sc h, H irkan-Wald, Parake nt Umg. , 2 0-18 0 m, D ol in le g." 6 e xs. , "25.5. 1999 Aze rba jdz ha n: Taly sc h, Suwant (Platea u) , 176 0 m, Dolin le g."; 3 0 e xs. , " Aze rba jdz ha n: Taly sc h, Zuwant platea u, 15 00 m, Go smol ja n e nv. [ca. 38.4 0N 48.2 1E ], 27.5.2 000, le g. Dolin ( AT 5)"; 1 #, 3 $$, "3 1.5. 1999 Aze rba jdz ha n: Taly sc h, Astara Umg. [= e nvirons; ca. 38.28N 48.52E ], Bac h und Fl uss, Dolin le g." ( a l l i n N MW) ; 1 $, "Paleke sh e nv. [ca. 38.3 0N 48.37E ], 13.- 15. V. 1992" , " Ase rbaid sc ha n, A. Sc ha mae v" (N MB) ; 1 $, " Aze rbai ja n Taly sc h Mt s. , km 14-18 , Rte Le nkora n-Lerik, 3 0. IV.-9. V.2 001, T. Lack ne r" , "Coll. He ndr ich, Be rlin" ; 1 $, " A z e rbai j a n T a l y s c h M t s . , L e rik m o un t a ins , 1 7 00m, 1 .- 3. V.2 001, T. Lack ne r" , "Coll. He ndr ich, Be rlin" ( a l l i n CLH); Iran : 1 #, " Iran: Gila n, 2 0 k m E Khalk hбl , 19 00 m, 13.- 16. V.2 000, le g. J. Re jsek" ( SMN S) ; 1 #, " Iran , ce ntr. Gila n, Ma sule h, (37.1 0N/48 .58E) , 55 k m WSW Ra sht , J. Re jsek 15.- 17.4.[ 19]99"; 1 #, " Iran , NE Maza ndara n, Nahar Khoran, (36 . 45 N/54 .28 E) , 6 k m south Gor ga n, J. Re jsek 15.- 17.4.[ 19]99"; 1 $, " Iran , W Maza ndara n, 10 k m N W Ha san Ke if, (36.36N/ 5 1. 10E) , J . Re jsek 2 0. -21.4.[1 9 ]99" ( a l l in C J S ) ; 1 #, "7.8. 1995 Iran , W. Aze rbay ja n, 75 k m N Sa ghez, wetla nd (t15) , 1400 m, El mi le g. ( # 165 0)" ; 1 #, "2.9. 1998 Iran , Ard eb il , 53 k m W Assa le m, strea m, 198 0 m, El mi le g. ( #2 13 1)"; 1 #, " 18.4.2 001 Iran , Fars, 17 k m SE Sep ida n (= Ardakan) , She sh Pir" , "ditc h with r unning wate r , El mi & Fe ry l e g." ( a l l i n CH F). Tu nisia : 1 #, " Tune sie n, 28.3.[ 19]9 1, Toujane , Q uelle [= sprin g], le g. We walka (T3)" [hw We walka] (N MW). 9 e xs. , "7.4.[ 19]92 Tune sie n, NE Tunis, Ca rthago, R innsal [= strea mlet ], Fe ry le g."; 2 e xs. , "8.4.[ 19]92 Tune sie n, S Tunis, El Zahra, Tьmpel [= pond] , Fe ry le g." ( a l l i n CH F). Algeria: 1 #, "Edo ugh, juin [18]85", coll. Bedel ( MNHN). Russia ( f i rst reco rd fro m Asia n R ussia -- the Ural s): 1 #, 3 $$, "C helyab. obl . [= C helya binskaya obla st = C helyab insk area ], Il me nsk. zap. [= Il menskiy zapo ved nik = Il me nskiy Nat ur e Re se rv e ], 22. VI. 194 1" [Cy rillic] (DE UM).

labels, but with the female sex symbol or without any such symbol (all in CGW). 1 #, 1 $ (matt), "Ost-Turkestan, Bagratsch-Kul" [hw Wewalka], "Locus typicus aus der Typenserie [= locus typicus of the type series], G. Wewalka" [hw Wewalka in part], "Hydroporus goldschmidti Gschw., det. G. Wewalka [19]72" [hw Wewalka in part]; specimens with head brownish, but not as light as in the types of H. recidivus (MRTO). Inexact localities: 1 #, "Turkestan"; 1 $ (smooth) "Samarkand" (all in NMW). Both specimens are rather old and have the head as brownish as in the types of H. recidivus. Samarkand is certainly used in the sense of the 19th century Russian Empire area of Samarkand which enclosed today's southern Uzbekistan and northern Tajikistan (see Schьtze & Kleinfeld [1997: 80]). Definitely, this specimen has not been collected in China.

DESCRIPTIVE NOTES. Hydroporus goldschmidti resembles externally Hydroporus planus (Fabricius, 1781), and shares with this species the reticulated last abdominal sternum. It has, however, the sides of the pronotum -- at least the lateral rim -- in most specimens brownish, the elytra of more reddish brown, and the total length on average smaller. The lighter spots behind the base of elytra are often rather distinct, whilst the elytral base of H. planus is only rarely considerably lightened. Body length and shape are strongly variable, as well as the extension of the pronotal reticulation, and cannot be used to separate the two taxa as it has been done by Wewalka [1992] who constructed, however, his key on the base of only a few examined specimens. On the other hand, H. goldschmidti and H. planus can be easily separated by the shape of the male genitalia (cf. Figs 7-9 and 10-12). MEASUREMENTS. TL: 3.3-4.2 mm, MW: 1.7-2.1 mm, TL/MW: 1.9-2.1 (lectotype of H. goldschmidti: TL: 3.9 mm, MW: 2.0 mm; lectotype of H. recidivus: TL: 4.0 mm, MW: 2.1 mm). DISCUSSION. The area of distribution of the original type series of H. goldschmidti is in the Tian Shan mountain range, and reaches from eastern Kyrgyzstan to north-western China (Xinjian). The locus typicus of H. recidivus is within H. goldschmidti's distribution area. Furthermore, one paralectotype of H. recidivus is very similar to H. goldschmidti; in particular, the penis coincides absolutely with that of H. goldschmidti (see also Wewalka [1992: 58]). In addition, we have not found significant differences in the female genitalia of the two taxa. The only differences between H. recidivus and H. goldschmidti are the colour of the head and the pronotum: head in large parts reddish brown in H. recidivus, only at anterior and posterior margins (elsewhere dark brown) in H. goldschmidti; sides of pronotum lighter brown to rather large extent in H. recidivus, only rim of pronotum lighter brown in most H. goldschmidti. All these observations induced us to treat H. recidivus as a junior subjective synonym of H. goldschmidti. DISTRIBUTION. China (Xinjian), Kyrgyzstan, Uzbekistan.

Hydroporus pubescens (Gyllenhal, 1808)
Gyllenhal, 1808: 536 (Hyphydrus pubescens) Type locality -- Sweden = Hydroporus distinguendus var. estrellensis Schaufuss 1882: 559 (Type locality -- Portugal, Serra da Estrкla)* Stephens, 1828: 61; Zimmermann, 1920: 100 (including synonymy of H. distinguendus var. estrellensis); Wewalka, 1992: 59; Nilsson & Holmen, 1995: 50; Nilsson, 2001: 158, 2003: 64. MATERIAL. Syntypes of Hyphydrus pubescens not studied. Type material: Lectotype of Hydroporus distinguendus var. estrellensis (by present designation): #, "Estrella, 18/ * Us ual ly Hydro po rus di st ing uen dus Des bro ch ers d es Lo ges, 1871 is treated as junior subjective synonym of Hydroporus jonicus L. Miller, 1862

DESCRIPTIVE NOTES. Specimens from the southern rt of the distribution area often have the elytra -- in rticular the basal part -- somewhat lighter yellowish brown. specimens from Iran these markings are particularly light d rather sharply delimited. DISTRIBUTION. Europe; Russia (at least southern European regions and the South Urals); North Africa except Egypt; Turkey; Cyprus; the Near East; Iran. pa pa In an

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae)

257

13

14

15

16

17

18

Figs. 13-18. Male genitalia of Hydroporus spp.: 13-15 -- H. limbatus; 16-18 -- H. brucki.; 13-15, 16-17 -- penis; 15, 18 -- paramere; 13, 16 -- ventral view; 14-15, 17-18 -- lateral view. Рис. 13-18. Гениталии самцов Hydroporus spp.: 13-15 -- H. limbatus; 16-18 -- H. brucki.; 13-15, 16-17 -- пенис; 15, 18 -- парамера; 13, 16 -- снизу; 14-15, 17-18 -- сбоку.

Hydroporus limbatus Aubй, 1838
Aubй, 1838: 292 Type locality -- Italy, Sardinia Z i mme r m a n n 1 9 31 : 1 3 4 ; G u i g n o t , 1 9 5 9 : 3 90 ; F r a n c i s c o l o, 1979: 336; Wewalka, 1992: 48; Nilsson, 2001: 157, 2003: 62. MATERIAL. Type material: Lectotype of Hydroporus limbatus (by present designation): $, "Limbatus" [hw Aubй], "Museum Paris, Coll. Aubй", "Lectotype, Hydroporus limbatus Aubй, 1838, Fery des. 2004" [red, printed]; specimen originally pinned, placed in the drawer behind a label "1975 Hydroporus Clairville, Limbatus Aubй" [hw Aubй] (MNHN). Notes: The lectotype has been glued onto a rectangular card by the senior author. It lacks the last six articles of the right antenna and the last tarsomere of the left hind leg. Behind the lectotype stands a female prepared in the same manner; it does not bear any label, except "Museum Paris, Coll. Aubй" (MNHN). This specimen is a Hydroporus lucasi Reiche, 1866, as can be easily seen by the reticulated last abdominal sternum and the dense and rugose punctation of its apical third. This specimen is not treated as paralectotype because it does not fit some details of the original description of H. limbatus, e.g., the rather coarse punctation of elytra and the absence of puncture lines. Aubй's description of H. limbatus suggests that the author had only a single specimen at his disposal. Since this is, however, not stated explicitly, the existence of syntypes instead of a holotype must be assumed (Recommendation 73F of the ICZN [1999]). Additional material examined: Portugal: 1 $, "Portugal mer. reg. Faro, Trafal /1/, 02.03.2004 Manti lgt., louka-sladkб voda" (CJH); 1 $, "7.11.[19]89 Portugal, Algarve, Vila Real, Tьmpel, Fery leg."; France: 11 exs., "4.2.[19]96 France, Camargue, Йtang du Charnier, Fery leg."; 1 #, 1 $, "Il [sic!] de Rй"; 1 $, "27.9.[19]88 France, Camargue, Aigues-Mortes, Lache, Fery leg."; 1 #, "31.3.[19]86 France, Herault, Agde, Wassergraben, Fery leg."; Italy: 1 $, "Sardinien 24.5./20.6.[19]65, Laloletta, Budberg"; 8 exs., "14.7.[19]81 Italia, Sardegna, Fertilia, Alghero, Tьmpel, Fery leg."; 2 exs., "Italie: Sicile - Catania, embouchure du Simeto, 11.04.[19]99, leg. G. Carron" (all in CHF). Unknown locality: 1 #, "Hydroporus exotique [?], velubinus mihi., h. [Dejean often used an "h." as abbreviation of "habitat", but here it was not followed by any locality.] D. Latreille"; a greyish and shiny label, text almost illegible, only perceptible if appropriately illuminated; the word "exotique" in red ink and of unknown hw, the rest in black ink, hw most probably that of Dejean; on the reverse "1221, 1222, 1223" [hw?], "Ex Musжeo Dejean" [printed], "limbatus" [hw Rйgimbart?], "D. Sharp Monogr." [printed], "Musйum Paris, ex coll. R. Oberthur, ex Wehncke";

specimen placed in the drawer behind a label "limbatus Aubй" [hw Wehncke] (MNHN). This specimen originally was mounted on a pin, and it has been glued onto a rectangular card by the senior author.

DESCRIPTIVE NOTES. This species is similar to H. brucki, but the punctation of the elytra is coarser and less dense; the puncture lines are only exceptionally and very weakly perceptible. The yellowish margin on sides of pronotum is somewhat narrower. In lateral view, the margin of the elytra is only weakly ascending towards humeral angle. According to Franciscolo [1979: 336] the total length is 4.0-5.0 mm; the examined specimens have 4.0-4.7 mm (lectotype 4.0 mm). The male genitalia are given in Figs 13-15; compare those of the externally similar H. planus in Figs 10-12. DISTRIBUTION. Italy, France, Spain, Portugal, Morocco, Algeria, Tunisia, and Libya. In Franciscolo [1979: 336], Corfu, Attica, and Crete are given additionally; these records, however, no doubt must be attributed to H. brucki.

Hydroporus brucki Wehncke, 1875
We h n ck e, 1 8 7 5 : 2 34 Type locality -- Greece, Thessaloniki ["Saloniki"]* Z i m m e r m a nn , 1 9 3 1 : 1 3 5 ; Z a i tz e v , 1 95 3 : 1 6 5 , 1 9 7 2 : 1 7 4; Wewalka, 1992: 48; Nilsson, 2001: 156, 2003: 60. MATERIAL. Type material: Lectotype of Hydroporus brucki (by present designation): $, "Brucki" [hw Wehncke], "Turcia, Salonichi" [hw Wehncke], "Hydroporus brucki Wenck., aus Coll. Oberthur" [hw Wewalka], "Musйum Paris, ex coll. R. Oberthur, ex Wehncke", "Lectotype, Hydroporus brucki Wehncke, 1875, Fery des. 2004" [red, printed]; specimen originally glued onto a small triangular tip, placed in the drawer behind a label "Brucki mihi" [hw Wehncke] (MNHN). Paralectotype: 1 $, a very small quadratic label without any text, "Musйum Paris, ex coll. R. Oberthur, ex Wehncke", and the respective red paralectotype label; specimen placed behind the lectotype, prepared in a similar manner as the lectotype (MNHN). Notes: Both specimens have been glued onto larger rectangular cards by the senior author. The lectotype lacks the left protarsus, three right protarsomeres, left mesotibia and mesotarsus, four right mesotarsomeres, left metatarsus, and the last right metatarsomere. The paralectotype lacks three left mesotarsomeres and the right metatarsus. * In 18 75 , Th es salo ni ki b el on ged st il l to t he Turkish (Ot toman or Osman) Empire

258

H. Fery, P.N. Petrov
21.IV.1992, leg. Schawaller"; 23 exs., "Appennino Romagnolo, Passo d. Muraglione, m 850 21.2.1993, leg. F. Pederzani" (all in CHF).

Additional material examined: Greece: 1 #, "Griechenland 1991, Ebrosebene (2), leg. Schцdel 16.V." (NMW); 1 #, "Corfu, Paganetti"; 1 #, "Kephallenia 1905, Argostolion, O. Leonhard"; 4 exs., "Kephallenia, Paganetti"; 1 #, "GR (38), Makedonia (CHA), Chalkidiki, Chlomуn Oros, Tal am Sьdhang QU 32, 16. 650 m, 29.05.[19]91., leg. CIT & GER"; 1 ex., "Olympos, Graecia, 2100 m, 4.6.[19]37, Coll. Bartoт"; 1 ex., "Gr. Ins. Lesbos, 4 km W. Agiassos, leg. Malicky 26.5.[19]75"; 1 ex., "GR 95-21: Insel Lesbos, SE, 12 km W Mitilнni, FluЯmьndung [= river mouth], Tьmpel [= pond]; leg. D. Grimm. 17.- 21.V.1995"; 10 exs., "GR Korinth 12.6.[19]86, Kato Alephori, leg. H. Hebauer" (all in CHF).

DESCRIPTIVE NOTES. The punctation of the elytra is rather dense and fine, approaching almost that of Hydroporus marginatus (Duftschmid, 1805). Other characteristic features are the broadly brownish sides of the pronotum and smooth abdominal sterna (in contrast to H. planus), the distinct puncture lines on the elytra, and the margin of the elytra distinctly ascending towards humeral angle (in contrast to H. limbatus). According to Zimmermann [1931: 135], its total length is 4.0- 4.3 mm, yet most of the examined specimens are distinctly longer: 4.5-4.7 mm, only one of these has 4.3 mm, another one 4.4 mm; the total length of the two types is 4.5 mm. The male genitalia are given in Figs 16-18. DISTRIBUTION. Greece, Turkey, Cyprus, and Lebanon.

DESCRIPTIVE NOTES. Hydroporus analis and H. decipiens are the only members of the planus-group that have a strongly rugose abdomen. Their upper surface is similar to that of the lighter coloured H. pubescens specimens. The male genitalia of H. analis are shown in Figs 19-21. The total length of the specimens studied is 3.4-3.8 mm (lectotype 3.7 mm). Franciscolo [1979: 331] gives 3.3-3.5 mm, Zimmermann [1931: 134] 3.5-3.75 mm. DISTRIBUTION. Italy, France (Corsica); records from Austria, Greece, Algeria, and Tunisia [Franciscolo, 1979: 341] should be confirmed; those from North Africa possibly must be attributed to H. decipiens.

Hydroporus decipiens Sharp, 1878
Sharp, 1878: 113 Type locality -- Spain, Sierra de Guadarrama Zimmermann, 1931: 134 (analis decipiens); Franciscolo, 1979: 340 (analis [= decipiens]); Fery & Fresneda, 1988: 342 (analis decipiens); Ribera et al., 1999: 57; Nilsson, 2001: 156, 2003: 60. MATERIAL. Type material: Lectotype of Hydroporus decipiens (by present designation): $, "Hydroporus decipiens Sharp, Type D.S." [on glue card, hw Sharp], "Type" [printed, round label with red margin, most probably mounted by J. Balfour-Browne], "Spain" [printed], "Sharp Coll, 1905 - 313" [printed], "Guadarrama" [hw Sharp], "Type 410, Hydroporus decipiens, Europe" [hw Sharp], "Lectotype, Hydroporus decipiens Sharp, 1878, Fery des. 2005" [red, printed] (BMNH). Paralectotype: 1 $, "Hydroporus decipiens Sharp, Ind. typ. D.S., Guadarrama" [on glue card, hw Sharp; the word "decipiens" on a small piece of paper glued onto the card], "Spain" [printed], "Spain" [printed], "Sharp Coll, 1905 - 313." [printed], "Syntype" [printed, round label with light blue margin, most probably mounted by J. Balfour-Browne], and the respective red paralectotype label (BMNH). Notes: The lectotype lacks the last three ar tic les of the r ight a ntenna. R. Bo oth (BMNH ) k ind ly communicated: "There is a label in the collection, probably in Garth Foster's handwriting, indicating that H. decipiens is a valid species and not a synonym of H. analis." According to the original description, Sharp has studied at least one further specimen from Monchique (southern Portugal) collected by Camille van Volxem. In the BMNH, however, no further material could be localised. Additional material examined: Portugal: 1 #, 2 ##, "9.8.[19]84 Portugal, Serra Estrella, b. Manteigas, Bach, 600 m, Fery leg."; 8 exs., "4.8.[19]89 Portugal, Serra Estrela, Umg. [= environs] Manteigas, kl. Bach, Fery leg."; 1 #, "22.7.1991 (P) Guarda-distr., Serra da Estrкla, N Manteigas, ca. 1100 m, Rinnsal, Fery leg."; 7 exs., "9.8.[19]84 Portugal, Serra Estrella, sьdl. Guarda, Rinnsal, Fery leg."; 9 exs., "14.8.[19]85 Portugal, Serra Estrela, Penhas Douradas, Bach, 1150 m, Fery leg."; 8 exs., "14.8.[19]85 Portugal, Serra Estrela, Penhas Douradas, Bach, 1000 m, Fery leg."; 9 exs., "21.7.[19]87 Portugal, Serra da Estrкla, Umg. [= environs] Torre, ca. 1900 m, Moortьmpel, Fery leg."; 1 ex., "30.1.[19]94 Portugal, Minho, Serra de Arga, Rinnsal, 800 m, Fery leg."; 1 $, "20.7.[19]87 Portugal, Odemira, Cercal, Rinnsal, Fery leg." (all in CHF). Spain: 1 #, 1 $, "Esp. Salamanca prov., La Alberca 1000 m, 5.6.2002 R. Sejkora lgt." (CJH); 15 exs., "23.7.[19]87 Espaсa, Prov. Leon, Astorga (sьdl.), Bach/Tьmpel, Fery leg."; 1 #, "6.7.[19]92 Espaсa, Prov. Leуn, Noceda, NE Ponferrada, Bach, Fery leg."; 10 exs., "22.7.[19]87 Espaсa, Prov. Zamora, Puente de Sanabria, Bach, Fery leg."; 5 exs., "21.3.[19]89 Espaсa, Prov. Zamora, Justel, цstl. Sanabria, Bach, Fery leg."; 12 exs., "3.11.[19]89 Espaсa, Prov. Zamora, O. [= E of] P. d. Sanabria, Bach, Justel, Fery leg."; 1 #, "4.4.1999 Spain, Burgos, ca. 3 km W Soncillo, ca. 30 km E Reinosa, ponds on meadow, Fery leg."; 1 #, "6.7.[19]88 Espagna, S. d. Gredos, Avila, Tormellas, Bach, Fery leg."; 2 exs., "9.7.[19]88 Espaсa, Prov. Avila, S. d. Gredos, Gredos, 1800 m, Quelltьmpel, Fery leg."; 30 exs., "3.8.[19]89

Hydroporus analis Aubй, 1838
Aubй, 1838: 294 Type locality -- Italy, Sardinia Zimmermann, 1931: 133; Guignot, 1959: 394; Wewalka, 1992: 47; Nilsson, 2001: 156, 2003: 60. MATERIAL. Type material: Lectotype of Hydroporus analis (by present designation): #, "Sardaigne" [a small round label, hw Aubй], "Museum Paris, Coll. Aubй", "Lectotype, Hydroporus analis Aubй, 1838, Fery des. 2004" [red, printed]; specimen placed in the drawer above a label "1976 Hydroporus Clairville, Analis Aubй" [hw Aubй?] (MNHN). Notes: The lectotype was originally pinned; it has been glued onto a rectangular card by the senior author. It lacks two left protarsomeres and the right hind leg. In the original description Aubй wrote "Il a йtй trouvй en Sardaigne et fait partie de la collection de M. Chevrolat". We have not been able to locate syntypes of H. analis in the collections of the IRSN or BMNH where parts of the collection of Chevrolat are said to have been preserved (see Horn et al. [1990: 71)]. However, we have no doubt that the specimen from the collection Aubй has been used by the author for his description. We assume that Aubй has kept the specimen for his collection after the study of Chevrolat's material. Additional material examined: France, Corsica: 1 #, "8.8.[19]74 Serriera, b. Porto (Corse), fast ausgetrock. FluЯ, Fery leg."; 1 ex., "2.4.[19]80 Korsika, Serriera, FluЯ am Strand, Fery leg."; 1 #, "3.4.[19]81 Korsika, Serriera, Lache am Strand, Fery leg."; 3 exs., "10.4.[19]81 Corse, Serriera (Porto), Tьmpel, Fery leg."; 12 exs., "10.4.[19]81 Corse, Serriera, Tьmpel, Fery leg."; 1 ex., "18.7.[19]82 Corse, Serriera, Bach, Fery leg."; 2 exs., "7.4.[19]80 Korsika, Francardo, Waldbach, Fery leg."; 1 ex., "7.4.[19]80 Corse, Casamozza, ehem. Kiesgrube, Fery leg."; 3 exs., "13.4.[19]81 Corse, Casamozza, Tьmpel, Fery leg."; 1 $, "13.4.[19]81 Corse, b. Barchetta, FluЯ Golo, Fery leg."; 1 #, "24.7.[19]81 Corse, Sc. di St. Regina, FluЯ Golo, Fery leg."; 1 #, "28.7.[19]81 Corse, Nдhe Fango, Bach, Fery leg."; 25 exs., "13.7.[19]83 Corse, Ponte Leccia, Stranciacone, Fery leg."; 2 exs., "19.4.[19]79 Korsika, Cargиse, Wiesenbach, Fery leg."; 1 ex., "26.7.[19]82 Corse, Sьdlich Cargиse, Lache, Fery leg."; 1 ex., "1.8.[19]74 Soccia (Corse), Tьmpel, Fery leg."; 8 exs., "8.4.[19]80 Korsika, Nцrdl. Sagone, FluЯ, Fery leg." (all in CHF). Italy: 2 exs., "I. Sardinien (201), цstl. Badde Suelzu, leg. Malicky 19 .10.[19 ]81"; 14 e xs., "18.7 .[19]81 Sa rd inien, Vil la gra nd e, Bach, Fery leg."; 8 exs., "18.7.[19]81 Sardinien, Villagrande, Tьmpel, Fery leg."; 2 exs., "Sardinien: NU, Stat. Arzana 800 m,

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae)

259

19

20

21

22

23

24

25
Figs zimmerm vie w. Рис. zimmerm

26

27

19-27. Male genitalia of Hydroporus spp.: 19-21 -- Hydroporus analis; 22-24 -- H. decipiens; 25-27 -- H. anni; 19-20, 22-23, 25-26 -- penis, 21, 24, 27 -- paramere; 19, 22, 25 -- ventral view; 20-21, 23-24, 26-27 -- lateral 19-27. Гениталии самцов Hydroporus spp.: 19-21 -- Hydroporus analis; 22-24 -- H. decipiens; 25-27 -- H. anni; 19-20, 22-23, 25-26 -- пенис, 21, 24, 27 -- парамера; 19, 22, 25 -- снизу; 20-21, 23-24, 26-27 -- сбоку.

Espaсa, Prov. Avila, Sierra de Gredos, Gredos, Tьmpel, 1800 m, Fery leg."; 3 exs., "9.7.[19]95 Espaсa, Prov. Avila, Sierra de Gredos, Gredos, Rinnsal, 1800 m, Fery leg."; 1 #, "22.5.[19]90 Espaсa, Prov. Guadalajara, S. d. Alto Rey, Bustares, Bach, Wiese, Fery leg."; 2 exs., "16.6.[19]90 Espaсa, Prov. Guadalajara, S. d. Alto Rey, Bustares, Rinnsal, Fery leg."; 1 #, "19.2.[19]90 Espaсa, Prov. Guadalajara, Anquela del Ducado, salzh. Lache [= salty puddle], Fery leg."; 8 exs., "24.7.[19]87 Espaсa, Prov. Madrid, Sierra de Guadarrama, Lozoya, Bach, 1300 m, Fery leg."; 2 exs., "1.7.[19]92 (E) Toledo, SW Los Yйbenes, Bachtьmpel, Fery leg."; 2 exs., "27.5.[19]90 Espaсa, Prov. Cбceres, S Coria, Caсaveral, Bachtьmpel, Fery leg."; 1 #, "2/iv/1996 Spain Cбcares [sic!], Arroyo de las Canteras, 5 km NE of Trujillo, granite stream, D.T. Bilton leg."; 1 $, "8.7.[19]95 (E) Palencia, Ledigos, E Sahagun, Rio Cuenza, Bach, Fery leg."; 1 ex., "27.5.[19]90 Espaсa, Prov. Ciudad Real, W Retuerta del Bullaque, Rinnsal, Fery leg."; 8 exs., "27. 5. [19]90 Espa сa , P ro v. C iudad Real , W Ret ue r ta del Bullaque, Bachtьmpel, Fery leg."; 1 ex., "4.8.[19]85 Espagna, Sierra Nevada, Laujar, Paterna del Rio, Rio de Alcolea, ca. 100 m, Fery leg."; 8 exs., "19.7.[19]87 Espaсa, Prov. Cadiz, nцrdl. Jimena d. l. Frontera, ZufluЯ d. Rio Hozgarganta, Fery leg." (all in CHF).

4.0 mm, paralectotype 3.9 mm). The male genitalia are figured for the first time (Figs 22-24). This taxon has been treated by most authors of the last century either as a synonym or a variety of H. analis. Ribera et al. [1999: 57] were the firsts who re-instated H. decipiens as a valid species, but did not give any reason for their procedure. DISTRIBUTION. Portugal and Spain; see also the notes under the distribution of H. analis.

Hydroporus crinitisternus Shaverdo & Fery, 2001
Shaverdo & Fery, 2001: 34 Typ e lo cality -- Kazakhs ta n , SW Dzhambul [= Auliye-Ata] Nilsson, 2001: 156. MATERIAL. Kazakhstan: Tarbagatay Mts., 4 km NE P 1100 m, 22.VI.2001, leg. W. S Talassk y Alatau rang e, 40 km

DESCRIPTIVE NOTES. Externally similar to in particular, with respect to the rugose surface of Yellowish spots on elytra more prominent. On av ger, total length of specimens studied 3.6-4.2 mm

H. analis, abdomen. erage lon(lectotype

Distribution: This new record completes well the known distribution of this rather poorly known species, which so far has only been recorded from south-eastern Kazakhstan and western Mongolia. The species seems to prefer the transition between the plain and mountainous regions. It should be noted that the Mongolian locality and the Kazakh ones are well connected by the Irtysh valley.

1 #, 1 $, "E Kazakhstan, S etrovskoe [ca. 47.03N 82.18E], chawaller" (SMNS).

260 Hydroporus tessellatus-group

H. Fery, P.N. Petrov
DISCUSSION. The synonymy of H. habelmanni has been treated in different ways (cf. list of references above). Gozis [1914: 161] treated the taxon as a variety of H. tessellatus, and some pages further [Gozis 1915: 178] as a variety of H. pubescens, however, remarking in a footnote that the majority of authors of that time preferred the former view. B ur me is te r [19 39 : 22 6] c all ed t he t axo n a "f or m" of H. pubescens, but not expressly giving it infrasubspecific rank (cf. ICZN [1999], Article 45.6, which might be applied here analogously, although such cases are not treated by the Code). To our knowledge, Franciscolo [1979: 334] was the first who gave it expressly infrasubspecific rank and called it "ab. habelmanni". However, Wewalka [1986: 278] and Angelini [1988: 47] still listed it as a subspecies of H. pubescens, whereas B. Guйorgiev [2004: 398] was the first who provided this taxon simply as a synonym of H. pubescens. As shown above, populations of H. pubescens from the Mediterranean, which usually are longer, broader, more vaulted, and have the base and the margins of elytra more yellowish than more northern ones, in the last decades have been treated either as a subspecies or as a colour variety of this species and called "habelmanni". The designation of the lectotype of H. habelmanni is at the same time a synonymisation of this name with H. tessellatus. Thus, these southern populations of H. pubescens lose their name, but, at present, we do not see the necessity to give them another one.

Hydroporus tessellatus (Drapiez, 1819)
Drapiez, 1819: 43 (Dytiscus tessellatus) Type locality -- Belgium, Moosel = Hyd ropo r us ha b elma n ni We hn ck e , 1 8 76 : 7 6 sy n . n. ( Type localit y -- Dalmatia*) He y d e n , 1883 : 30 ( hab el m anni ), 189 1 : 62 ( pub esce ns v a r. h a b e lm a n ni), 1 9 06 : 1 16 (t e s s el a t us v ar. h a b e l m a n ni ); S e i d li t z , 1 8 8 7 : 7 2 ( p u b es ce ns v ar . h a b elm a n ni ) ; J a co b s o n , 1 9 0 8: 42 5 ( p u b es c e n s h a b el m an ni ) ; Z i mme r ma nn, 1 92 0 : 1 00 ( p ub es c e ns h a b elm a n ni ); B u r mei s ter, 1 9 39 : 2 2 6 ( pu b esce n s f. h a b el m an ni ) ; F r a n ci s c o l o, 1 9 79 : 3 34 ( p u b es c e n s a b . h a b e lm a n ni ) ; W e wa l k a , 1 98 6 : 2 78 ( p u b e s ce n s ha b el m an ni ) ; A ngelini, 1988: 47 (pubescens habelmanni); Balke & Fery 1993: 93 ( n e o t y p e d esi gn at i on of H. hab e lm anni ); N i lss on , 2 001: 16 6, 2 0 03 : 65. MATERIAL. Type material: Neotype of Hydroporus tessellatus designated by Balke & Fery [1993: 93], stored in the IRSN. Lectotype of Hydroporus habelmanni (by present designation): $, a small quadratic label without text, "Dalmatia" [hw Wehncke], "ex Wehncke, Musйum Paris, ex coll. R. Oberthur", "Lectotype, Hydroporus habelmanni Wehncke, 1876, Fery des. 2004" [red, printed]; glued onto a small triangular tip, placed in the drawer behind a label "Habelmanni mihi" [hw Wehncke] (MNHN). Paralectotype: 1 #, "Steinheil" [? nearly illegible, hw Wehncke], "Elba" [hw Wehncke], "ex Wehncke, Musйum Paris, ex coll. R. Oberthur", and a respective red paralectotype label; placed behind the lectotype and prepared in the same manner (MNHN). Note. The lectotype is an immature Hydroporus tessellatus (Drapiez, 1819) with a total length of 3.5 mm; it lacks the left hind tarsus. Both types have been re-glued by the senior author onto larger rectangular cards.

Hydroporus zimmermanni G.[J.] Mьller, 1926
Mьller, 1926: 288 Type locality -- Slovenia, Pivka [= St. Peter = San Pietro del Carso], ca. 30 km E Trieste [Italy] = Hydroporus bicolor J.[G.] Mьl ler 1933: 202 (Typ e locality Sl ov en i a , Pi vk a) Mьller, 1928: 260, 1933: 201; Zimmermann, 1931: 127; Gs ch we n d tner, 1939: 34 (bicolor); Zaitzev, 1953: 167 (in cluding bicolor), 1972: 176 (including bicolor); V.B. Guйorguiev, 1960: 27, 28 (bicolor), 1965: 492 (including bicolor), 1971: 13 (including synonymy of H. bicolor); Nilsson, 2001: 167, 2003: 65. MATERIAL. Type material: Holotype of Hydroporus zimmermanni: #, "Carniolia, St. Peter, 9.4.[19]11", "Hydroporus Guernei Rйg." [hw Zimmermann], "Museo Civico di Trieste", "Typus" [red], "Zimmermanni Mьll., Tipo" [green, hw J.[G.] Mьller] (MCTR). Notes: Mьller's [1926: 288] original description of H. zimmermanni contains no hint on the number of syntypes studied. In a later publication [Mьller 1928: 260] the author provided "... the single exemplaire of H. Zimmermanni ..."; Zimmermann's [1931: 127] remark can be interpreted in the same way: "Type from St. Peter in Krain ...". Thus Mьller has a single exemplare of a new species. So, this specimen is a holotype by monotipy (Recommendations 72.4.1.1 and 73.1.2. of the ICZN [1999]). Lectotype of Hydroporus bicolor (by present designation): #, "Istr. [= Istria] St. Peter, 16.4.33, Springer" [printed, hw Springer (?) in part], "Petel. jezero" [hw Springer?], "Cotypus" [red], "Museo Civico di Trieste", "Lectotype, Hydroporus bicolor J.[G.] Mьller, 1933, des. H. Fery 2005" [red, printed] (MCTR); Paralectotypes: 1 $, first four labels as in the lectotype; 1 $, "S. Pietro, Petelinsko jezero" [hw Springer (?)], "Springer, 16.4.33" [hw Springer?], "Museo Civico di Trieste", "Typus" [red], "bicolor m., det. J. Mьller" [hw J.[G.] Mьller?] (all in MCTR). Each paralectotype is provided with the respective red label. Additional material examined: Slovenia: 1 ex., "S. Pietro, Petelinsko jezero" [hw Springer?], "Springer, 16.4.33" [hw Springer?], "Museo Civico di Trieste"; abdomen externally glued onto the card, genitalia missing; 2 exs., "Istr. St. Peter, 16.4.33, Springer" [date hw Springer?], "Petel. jezero" [hw Springer?], "Museo Civico di Trieste" (all in MCTR). 2 exs., "Galsko jezero [not found on any map], 7.V.1956", "Slovenia, leg. Pretner"; genitalia absent (MRTO). 4 exs., "Petelinsko Jezero", "7.V.1956.",

NOTES. It was not possible to identify the paralectotype of H. habelmanni undoubtedly. It is an extremely immature specimen of ca. 3.3 mm total length and resembles H. pubescens insofar as it has the elytra as well as the last abdominal sternum smooth between the punctures. On the other hand, the punctation on the apical third of this segment is rather sparse and not densely and almost rugosely punctate as it is typical for H. pubescens; the punctures are distinctly separated one from another and the smooth spaces between them clearly perceptible. The sculpture of this segment resembles much more that of H. limbatus, a species which has, however, usually a distinctly higher total length (see above). Additionally, due to the high immaturity it was not possible to identify the specimen by the shape of the aedeagus. A further reason to designate not the Elba specimen but the "Dalmatia" one as lectotype is that the latter fits better Wehncke's description with respect to some minute features which, however, shall not be explained here in detail. In the Wehncke collection, there stands a third specimen behind the lecto- and paralectotype which, however, is not treated as belonging to the syntype series because it is not mentioned in the original description: 1 ex., "Smyrna [= Izmir, Turkey]" [hw Wehncke], "ex Wehncke, Musйum Paris, ex coll. R. Oberthur" (MNHN). This specimen is -- like the lectotype -- an immature H. tessellatus. The designation of a specimen of H. tessellatus as lectotype is -- at first sight -- not in accordance with Recommendation 74A of the ICZN [1999], since in the last decades H. habelmanni was treated as a synonym of H. pubescens. It seems, however, to be absolutely necessary to base the designation of the lectotype on the specimen of clear identity and not on the one which might give rise to future doubts.
* This coincides more or less with today's Croatia

Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae)
"Slovenija, leg. Pretner" (NMW, CHF). 4 exs., "Petelinsko Jezero, Pivka, 21.4.1963", "YU Slovenien, leg. Pretner"; Serbia and Montenegro: 14 exs., "Crna Gora [= Montenegro], Durmitor, Skrcko Jezero, 1933.VII.13, leg. Dr. J. Fodor" (all in CHF).

261

DESCRIPTIVE NOTES. The types of H. bicolor deviate from other H. zimmermanni only by the almost uniformly light yellowish elytra. Thus, the contrast between these and the black pronotum is more prominent. The other separating characters given by Mьller [1933: 202] are not sufficient to create a new species. The male genitalia are figured for the first time (Figs 25-27). The total length of the specimens studied is 3.9-4.3 mm. DISTRIBUTION. According to Nilsson [2003: 65], this species is distributed in Slovenia, Croatia, Serbia and Montenegro, Bosnia Herzegovina, and Bulgaria; the latter, however, must be excluded due to the following observations: Hlisnikovskэ [1955: 96] recorded H. zimmermanni from Bulgaria. We have studied two females with the same collecting data: "Sozopol [ca. 42.25N 27.41E] Bulg. or. 16.V.[19]39, Hlisnikowski". These specimens have a total length of 4.4- 4.5 mm and are thus distinctly longer than H. zimmermanni. In addition, the punctation of the elytra is much denser and finer, the spaces between punctures are not reticulated. The specimens resemble Hydroporus inscitus Sharp, 1882, or even Hydroporus multiguttatus Rйgimbart, 1887 (Hydroporus marginatus-group), but they are definitely not H. zimmermanni. For a solution of this problem, the study of males is indispensable.

Hydroporus marginatus-group Hydroporus marginatus (Duftschmid, 1805)
Duftschmid 1805: 269 (Dytiscus marginatus) Type locality -- Austria, Kleinmьnchen, near Linz S t e p h e n s , 18 2 8 : 56 ; F ra n cis co l o , 19 7 9 : 32 7 ; Nil s s o n , 20 0 1: 160, 2003: 62. Type material: According to Gusenleitner [1984], all Duftschmid's types must be treated as not to be found. Material examined: Turkey: 1 #, "Dumlu-daрl. bei Akdagkцy, 2300-2900 m, 21.V.1989", "Anatolia or. [= oriental], Heinz leg." (NMB). Kazakhstan (first record): 1 #, "NKazakhstan Reg., Markakol, Alekseyevka, 1700 m 27.VI.1987, V. Dolin" (NMB). Iran (first record): 4 exs., "15.8.1998 Iran, Kohkiluyeh & Boyer Ahmad, 7 km SW Yasuj, Elmi & Fery leg.", "Cheshmeh Sar-e-Ab Taveh, brook and pools (with stagnant water) (#2106)" (CHF).

ACKNOWLEDGEMENTS. We are indebted to all the colleagues and the curators who kindly made material available for our studies. We thank A.N. Nilsson (Umeе, Sweden), who contributed with several helpful discussions. We also thank D. Telnov (Riga, Latvia) for help with the transportation of some specimens.

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